''Pendraig'' (meaning "chief dragon" in
Middle Welsh
Middle Welsh ( cy, Cymraeg Canol, wlm, Kymraec) is the label attached to the Welsh language of the 12th to 15th centuries, of which much more remains than for any earlier period. This form of Welsh developed directly from Old Welsh ( cy, Hen ...
) is a genus of
coelophysoid
Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from
South Wales
South Wales ( cy, De Cymru) is a loosely defined region of Wales bordered by England to the east and mid Wales to the north. Generally considered to include the historic counties of Glamorgan and Monmouthshire, south Wales extends westwards ...
. It contains one species, ''Pendraig milnerae'', named after
Angela Milner
Angela Cheryl Milner (3 October 1947 – 13 August 2021) was a British paleontologist who, in 1986 alongside Alan Charig, described the dinosaur '' Baryonyx''.
Early life
Milner was born Angela Girven in Gosforth, daughter of Cyril and L ...
. The specimen was discovered in the
Pant-y-Ffynnon quarry
Pant-y-Ffynnon Quarry is a stone quarry in the Vale of Glamorgan, Wales, around 3 kilometers east of Cowbridge. It contains fissure fill deposits dating to the Late Triassic (Rhaetian), hosted within karsts of Carboniferous aged limestone, pri ...
. In life it would have measured in length.
History
The holotype of ''Pendraig'' were found in the
Pant-y-ffynnon Quarry
Pant-y-Ffynnon Quarry is a stone quarry in the Vale of Glamorgan, Wales, around 3 kilometers east of Cowbridge. It contains fissure fill deposits dating to the Late Triassic (Rhaetian), hosted within karsts of Carboniferous aged limestone, pri ...
in Wales in 1952 by Kermack and Robinson along with the holotypes of ''
Pantydraco
''Pantydraco'' (where "panty-" is short for Pant-y-ffynnon, signifying ''hollow of the spring/well'' in Welsh, referring to the quarry at Bonvilston in South Wales where it was found) was a genus of basal sauropodomorph dinosaur from the Late Tria ...
'' and ''
Terrestrisuchus
''Terrestrisuchus'' is an extinct genus of very small early crocodylomorph that was about long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by t ...
'', and were subsequently lost in the collections of the
Natural History Museum, London
The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum ...
. The fossils were originally thought to belong to a "
coelurosaur
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
" (in the outdated sense of the word) and even subsequently classified as a species of "''Syntarsus''" (now ''
Megapnosaurus
''Megapnosaurus'' (meaning "big dead lizard", from Greek μεγα = "big", 'απνοος = "not breathing", "dead", σαυρος = "lizard") is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago dur ...
'' or ''
Coelophysis
''Coelophysis'' ( traditionally; or , as heard more commonly in recent decades) is an extinct genus of coelophysid theropod dinosaur that lived approximately 228 to 201.3 million years ago during the latter part of the Triassic Period fr ...
''). Recently Angela Milner and
Susannah Maidment Susannah Catherine Rose Maidment is a British palaeontologist at the Natural History Museum, London. She is internationally recognised for her research on ornithischian dinosaur evolution, and was awarded the 2016 Hodson Award of the Palaeontolo ...
rediscovered the fossils stored with some crocodile bones (likely ''
Terrestrisuchus
''Terrestrisuchus'' is an extinct genus of very small early crocodylomorph that was about long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by t ...
''). The fossils were named as representing a new genus of theropod in 2021.
Description

The holotype specimen consists of NHMUK PV R 37591, "An articulated vertebral series and pelvic girdle comprising the two posteriormost dorsal vertebrae missing most of the neural spines, the three anteriormost sacral vertebrae and a small fragment of the centrum of the fourth sacral, a complete left ilium, a largely complete left pubis missing the distal end, a left ischium missing most of the distal portion, a largely complete right pubis missing the distal end, and a right ischium missing most of the dorsal and distal portions" a left femur that is from the same individual was found disarticulated from the main block.
Classification
Spiekman ''et al.'' recover ''Pendraig'' as a
coelophysoid
Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
theropod, in a polytomy with ''Powellvenator'', ''Lucianovenator'', and the clade comprising "''Syntarsus''" ''kayentakatae'' and Coelophysidae. Their analysis is shown below:
Paleoecology

During the Late Triassic, the areas of southwestern Britain ''Pendraig'' was found and lived in consisted of a series of islands made from
Carboniferous Limestone
Carboniferous Limestone is a collective term for the succession of limestones occurring widely throughout Great Britain and Ireland that were deposited during the Dinantian Epoch of the Carboniferous Period. These rocks formed between 363 ...
that had been folded and pushed upwards. The environment would have been dry forests described as similar to the
Cayman Islands.
Other taxa known from the Pant-y-Ffynnon quarry that would have lived alongside ''Pendraig'' include the engimatic pseudosuchian ''
Aenigmaspina
''Aenigmaspina'' (from Latin ''aenigma'' and ''spina'', meaning "enigmatic spine") is an extinct genus of enigmatic pseudosuchian (=crurotarsan) archosaur from the Late Triassic of the United Kingdom. Its fossils are known from the Pant-y-ffynno ...
,'' the basal crocodylomorph ''
Terrestrisuchus
''Terrestrisuchus'' is an extinct genus of very small early crocodylomorph that was about long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by t ...
,'' the sauropodomorphs ''
Thecodontosaurus
''Thecodontosaurus'' ("socket-tooth lizard") is a genus of herbivorous basal sauropodomorph dinosaur that lived during the late Triassic period (Rhaetian age).
Its remains are known mostly from Triassic "fissure fillings" in South England. ''T ...
'' and ''
Pantydraco
''Pantydraco'' (where "panty-" is short for Pant-y-ffynnon, signifying ''hollow of the spring/well'' in Welsh, referring to the quarry at Bonvilston in South Wales where it was found) was a genus of basal sauropodomorph dinosaur from the Late Tria ...
'', the
rhynchocephalians ''
Clevosaurus
''Clevosaurus'' (meaning "Gloucester lizard") is an extinct genus of rhynchocephalian reptile from the Late Triassic and the Early Jurassic periods. Species of ''Clevosaurus'' were widespread across Pangaea, and have been found on all continents ...
'' and ''
Diphydontosaurus
''Diphydontosaurus'' is an extinct genus of rhynchocephalian reptile from the Late Triassic of England and Italy. This small animal was related to the living tuatara (''Sphenodon''). It may have grown to a length of . It is more derived than ' ...
,'' and the gliding reptile ''
Kuehneosaurus.''
Based on similar patterns on island ecosystems like Cayman Islands and
Aldabra
Aldabra is the world's second-largest coral atoll, lying south-east of the continent of Africa. It is part of the Aldabra Group of islands in the Indian Ocean that are part of the Outer Islands of the Seychelles, with a distance of 1,120 k ...
, it is likely there were large densities of these species due to a lack of known large predators and competition from other animals.
Due to its island habitat, the small size of ''Pendraig'' may result from
insular dwarfism
Insular dwarfism, a form of phyletic dwarfism, is the process and condition of large animals evolving or having a reduced body size when their population's range is limited to a small environment, primarily islands. This natural process is dist ...
, where animals on islands get smaller to conserve resources.
However, due to the specimen being sub-adult, it may or may not be the case.
See Also
*
Palaeontology of Wales
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (geology), epoch (roughly 11,700 years before present). It includes t ...
References
{{Taxonbar, from=Q108810741
Coelophysoids
Late Triassic dinosaurs of Europe
Fossils of Wales
Fossil taxa described in 2021
Paleontology in Wales