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''Pelargonium luridum'', locally called variable stork's bill, is a medium high,
tuber Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reproduc ...
ous
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
geophyte A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
, belonging to the Stork's bill family, with white to pink, slightly mirror symmetrical flowers in
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s on long unbranched stalks directly from the ground rosette that consists of few initially ovate, later pinnately incised or linear leaves, with blunt teeth around the margin. The variable stork's bill naturally occurs from South Africa to Angola, southern Congo and Tanzania.


Description

''Pelargonium luridum'' is a very variable species. It is an erect, perennial, herbaceous plant with up to long inflorescence stalks, with a woody tuber and with all leaves growing directly from the heart of the plant at ground level. The plants are covered in a layer of soft, about long hairs, and shorter hairs that are pressed to the surface of the plant. It also has glands. Most of the leaves appear after flowering. The
stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole (botany), petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part ...
s on both sides next to the base of the leaf stalks are linear or long triangular in shape and long. These are persistent and turn brown with age. The leaf stalks are usually , occasionally up to long. The
leaf blade A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
is oval in outline, usually long and wide, wedge- to
heart-shaped The heart symbol is an ideograph used to express the idea of the "heart" in its metaphorical or symbolic sense. Represented by an anatomically inaccurate shape, the heart symbol is often used to represent the center of emotion, including affecti ...
at its foot, variably incised, form shallowly lobed to multiply dissected, the later leaves often more complex. The segments at the margins of the leaf-outline, can be thread-thin to oval, entire of with teeth around the margin, and the tip pointy of rounded. The upper leaf surface may be sofly hairy or almost without, except for the margins and veins. Each plant may carry one to three
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s, consisting of a long peduncle with few to many small, pointy, simple membraneous bracts long and 1½–4 mm (0.06–0.16 in) wide, topped by a simple
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of usually between five and thirty, sometimes even fifty flowers. At the top of the peduncle is a whorl of small tapered bracts. The lower part of the stalk of the individual flower is mostly long. The
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s are deflected when the flower has opened, lanceolate to narrowly oval, 8½–15 mm (0.33–0.6 in) long and 1½–5 mm (0.06–0.20 in) wide, and are covered in dense felty hairs and glands. The flowers have a spur, that has merged with the upper part of the flower stalk of long. There are five
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s present, which are at first spreading but later somewhat reflexed, and are long inverted egg-shaped or lanceolate with the widest point towards the tip, usually long and wide. Their color varies from white to pale yellow, with a pink venation or entirely purple. The three anterior petals have a narrow basal half (a so-called
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or Arthro ...
) and a wider top half (or plate). The ten stamens are fused at their foot for 1¾–3½ mm (0.07–0.14 in). Usually seven of these stamens carry long, 1¼–1¾ mm (0.05–0.07 in) wide
anthers The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...
, and their filaments are long, the usually three infertile
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...
s are 3–5½ mm (0.12–0.22 in) long. The ovary is softly hairy, with a short
style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...
long, topped by five stigmas long. It develops into a
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
that grows to , exceptionally 6½ cm long. When ripe each of the five seeds comes of with a long and 1½–3 mm (0.06–0.12 in) wide part of the fruit called a
mericarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the ...
. Individual seeds within the mericarp are long and wide. The pale brown
seed coat In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be sown, which may include seed and husk or tuber. Seeds are the ...
has a very fine netted structure.


Taxonomy

Henry Cranke Andrews Henry Cranke Andrews (c. 1759 – 1835, floruit, fl. 1794 – 1830), was an English botanist, botanical artist and engraver. As he always published as Henry C. Andrews, and due to difficulty finding records, the C. was often referred to as Cha ...
was the first to describe this species in 1813 as ''Geranium luridum''. In 1820, Robert Sweet described a non-flowering plant that was grown in England by a mrs. Colville, from a rootstock sent from South Africa, calling it ''Pelargonium huraefolium''. After the plant had started flowering later, Sweet realised his species was in fact identical to that of Andrews, but remained of the opinion that it belonged to ''Pelargonium'', and so created the
new combination In Taxonomy (biology), biological taxonomy, a combinatio nova (abbreviated comb. nov. or n. comb.) refers to the formal renaming of an organism's scientific name when it is transferred to a different genus, reclassified within a different specie ...
''Pelargonium luridum'' in 1826. ''Polyactium aconitiphyllum'' was described based on a specimen from South Africa in 1835 by
Christian Friedrich Ecklon Christian Friedrich Ecklon (17 December 1795 – 1 December 1868) was a Denmark, Danish botany, botanical collector and apothecary. Ecklon is especially known for being an avid collector and researcher of plants in South Africa. Biography Ecklon ...
and
Karl Ludwig Philipp Zeyher Karl Ludwig Philipp Zeyher (2 August 1799 Dillenburg, Hessen, Holy Roman Empire – 13 December 1858 Cape Town), was a botanical and insect collector who collected extensively in Cape Colony. He was the author, with Christian Friedrich Ecklon, of ...
.
Ernst Gottlieb von Steudel Ernst Gottlieb von Steudel (30 May 1783 – 12 May 1856) was a German physician and an authority on grasses. Biography Ernst Gottlieb von Steudel was born at Esslingen am Neckar in Baden-Württemberg. He was educated at the University of Tübing ...
moved it in 1842 to ''Pelargonium'', creating the new combination ''Pelargonium aconitiphyllum''. Because the species is very variable, several later authors raised different morphs to their own species.
Ernst Heinrich Friedrich Meyer Ernst Heinrich Friedrich Meyer (1 January 1791 – 7 August 1858) was a German botanist and botanical historian. Born in the Electorate of Hanover, he lectured in Göttingen and in 1826 became a professor of botany at the University of Königsb ...
in 1843 named ''P. polymorphum'', but failed to adequately describe it. In 1860
William Henry Harvey William Henry Harvey, FRS FLS (5 February 1811 – 15 May 1866) was an Irish botanist and phycologist who specialised in algae. Biography Harvey was born at Summerville near Limerick, Ireland, in 1811, the youngest of 11 children. His father ...
distinguished ''P. flabellifolium'', and Harvey in cooperation with
Otto Wilhelm Sonder Otto Wilhelm Sonder (18 June 1812, Bad Oldesloe – 21 November 1881) was a German botanist and pharmacist. Life A native of Holstein, Sonder studied at Kiel University, where he sat pharmaceutical examinations in 1835, before becoming the pro ...
defined ''Pelargonium zeyheri'' in the same year.
Ignaz von Szyszyłowicz Ignaz von Szyszyłowicz (30 July 1857 – 17 February 1910) also known as Ignacy Szyszyłowicz was a Polish botanist born in Granica ( Sosnowiec). He contributed Part III.6 ''Caryocaraceae, Marcgraviaceae, Theaceae, Strasburgeriaceae'' to Engl ...
distinguished two other forms, naming them ''P. rehmannii'' and ''P. angulosum'' respectively in the year 1888.
Otto Kuntze Carl Ernst Otto Kuntze (23 June 1843 – 27 January 1907) was a German botanist. Biography Otto Kuntze was born in Leipzig. An apothecary in his early career, he published an essay entitled ''Pocket Fauna of Leipzig''. Between 1863 and 1866, he ...
moved ''flabellifolium'', ''rehmannii'' and ''zeyheri'' to a new, more narrowly defined genus ''Geraniospermum'', in 1891. In 1902,
Adolf Engler Heinrich Gustav Adolf Engler (25 March 1844 – 10 October 1930) was a German botanist. He is notable for his work on plant taxonomy and phytogeography, such as ''Die natürlichen Pflanzenfamilien'' (''The Natural Plant Families''), edited with K ...
describes ''P. heckmannianum''. All of these names are now considered
synonymous A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
to ''P. luridum''. ''P. luridum'' is the only species assigned to the subsection Magnistipulacea, which itself is part of the large section Polyactium.


Etymology

The species name ''luridum'' is from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
and means pale, sallow or smoky yellow.


Distribution and habitat

''Pelargonium luridum'' occurs naturally in Angola, Eswatini, Lesotho, Malawi, South-Africa (Cape Province, Natal, Orange Free State, Transvaal), Tanzania, Zambia, Zimbabwe. It can be found in grassland and open bush, is said to prefer locations subject to regular fires, and damp ground such as by streams.


References

{{Taxonbar, from=Q7213346 Plants described in 1813 luridum Flora of East Tropical Africa Flora of South Tropical Africa Flora of Southern Africa Taxa named by Henry Cranke Andrews