''Pectinodon'' is a
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
troodontid
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s from the end of the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
age of the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period (66
mya). It currently contains a single valid
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, ''Pectinodon bakkeri'' (sometimes classified as ''
Troodon bakkeri''), known only from teeth.
History of discovery

In 1982,
Kenneth Carpenter named a number of theropod teeth from the late Maastrichtian aged
Lance Formation
The Lance (Creek) Formation is a division of Late Cretaceous (dating to about 69–66 Ma) rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the lates ...
of
Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
as the
type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Pectinodon bakkeri''. The generic name is derived from the
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word ''pecten'', meaning "comb", and the Greek word ὀδών, ''odon'', meaning "tooth", in reference to the comb-like serrations on the rear edge of the teeth. The
specific name honors famed paleontologist
Robert Thomas Bakker.
[
The holotype, UCM 38445, consists of a 6.2 mm long adult tooth. The ]paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
s are three juvenile teeth.[
In 1985, Lev Nesov named a second species, ''Pectinodon asiamericanus'', based on specimen CCMGE 49/12176, a tooth from the Khodzhakul Formation of ]Uzbekistan
, image_flag = Flag of Uzbekistan.svg
, image_coat = Emblem of Uzbekistan.svg
, symbol_type = Emblem of Uzbekistan, Emblem
, national_anthem = "State Anthem of Uzbekistan, State Anthem of the Republ ...
that dates from the Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
age. This is today often considered a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''.
While historically considered synonymous with ''Troodon'' or more specifically the species '' Troodon formosus'', Philip Currie and colleagues (1990) noted that the ''P. bakkeri'' fossils from the Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
and Lance Formation might belong to different species. In 1991, George Olshevsky assigned the Lance formation fossils to the species ''Troodon bakkeri''.[Currie, P. (2005). "Theropods, including birds." in Currie and Koppelhus (eds). ''Dinosaur Provincial Park, a spectacular ecosystem revealed, Part Two, Flora and Fauna from the park.'' Indiana University Press, Bloomington. Pp 367–397.] In 2011, Zanno and colleagues reviewed the convoluted history of troodontid classification in Late Cretaceous North America. They followed Longrich (2008) in treating ''Pectinodon bakkeri'' as a valid genus and noted that it is likely the numerous Late Cretaceous specimens currently assigned to ''Troodon formosus'' almost certainly represent numerous new species, but that a more thorough review of the specimens is required.
In 2013, Currie and Derek Larson concluded that ''Pectinodon bakkeri'' was valid and its teeth could be found both in the Lance Formation and the coeval Hell Creek Formation. Some teeth from the older Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
could not be statistically differentiated from them, likely due to an insufficiently large sample, and were referred to ''cf. Pectinodon''.
Description
While ''Pectinodon'' is only known from teeth, its larger family Troodontidae is known from much more complete specimens. They were small, bird-like feathered bipedal maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, ...
ns with proportionally large eyes and brains. Like dromaeosaurids, they possessed a "sickle-claw" on the second toe of each foot. See the Troodontidae
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
article for more information.
''Pectinodon'''s eponymous teeth are generally small (< 1 cm crown height) labio-lingually compressed, exhibit basal constriction, and possess large, triangular, apically-oriented denticles on the distal edge. The holotype tooth has pitting on the lingual surface and no denticles on the mesial carina, though other specimens are known to have fine serrations near the base of the crown on the mesial carina.
Paleobiology
Despite having only teeth to infer behavior for ''Pectinodon'', they are very informative fossils. Several studies have been done on troodontid teeth, shedding light on their behavior and ecology. Their tooth morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
suggests they preferred soft prey items like invertebrates, small vertebrates, or carrion. Their dentition was not as well suited for high-stress feeding as in their cousins the dromaeosaurids and tyrannosaurids. Tooth morphology also led some researchers to speculate that troodontids were omnivorous
An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
. This hypothesis was reaffirmed in a study that used stable isotopes in tooth enamel to reconstruct paleo-ecologies.
See also
* Troodontidae
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
* Timeline of troodontid research
References
{{Taxonbar, from=Q4037858
Troodontidae
Dinosaur genera
Maastrichtian dinosaurs
Hell Creek Formation
Lance Formation
Taxa named by Kenneth Carpenter
Fossil taxa described in 1982
Dinosaurs of the United States