''Paratriacanthodes'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of marine

ray-finned fishes Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class (biology), class of Osteichthyes, bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Triacanthodidae, the spikefishes. These fishes are found found in the

Indian Indian or Indians may refer to: Associated with India * of or related to India ** Indian people ** Indian diaspora ** Languages of India ** Indian English, a dialect of the English language ** Indian cuisine Associated with indigenous peoples o ...

and

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...

Taxonomy

''Paratriacanthodes'' was first proposed as a

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

in 1934 by the American

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Henry Weed Fowler Henry Weed Fowler (March 23, 1878 – June 21, 1965) was an American zoologist born in Holmesburg, Philadelphia, Holmesburg, Pennsylvania. He studied at Stanford University under David Starr Jordan. He joined the Academy of Natural Sciences in Ph ...

when he described ''P. retrospinis'' as its only species, he also designated as the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

of the genus. When Fowler described ''P. retrospinis'' he gave its type locality as the

China Sea The China Seas consist of a series of marginal seas in the Western Pacific Ocean, around China. They are the major components signifying the transition from the continent of Asia to the Pacific Ocean.Pinxian Wang, Qianyu Li, Chun-Feng Li, ''Geol ...

in the vicinity of Taiwan at 21˚36'00''N, 117˚27'00''E,

Albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...

station D.5517 from a depth of . Later in 1935

George S. Myers George Sprague Myers (February 2, 1905 – November 4, 1985) was an American ichthyologist who spent most of his career at Stanford University. He served as the editor of ''Stanford Ichthyological Bulletin'' as well as president of the American S ...

added a second species ''P. herrei'' from the Philippines and in 1967 James C. Tyler added a third, ''P. abei'' from the South China Sea. In 1968 Tyler classified this genus in the

nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

of the family Triacanthodidae, the

Triacanthodinae Triacanthodinae is a subfamily of marine ray-finned fishes belonging to the family Triacanthodidae, the spikefishes. This subfamily comprises nine genera and a total of nineteen species and all, except one species, are found in the tropical an ...

. The 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' classifies the family Triacanthodidae in the

suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...

Triacanthoidei Triacanthoidei is a suborder of ray-finned fishes belonging to the order Tetraodontiformes, which includes the pufferfishes, triggerfishes and related taxa. These benthic fishes are mainly found in the Indian Ocean with some of the spikefishes fou ...

in the order Tetraodontiformes.

Species

There are currently 3 recognized species in this genus: * '' Paratriacanthodes abei'' Tyler, 1997 * '' Paratriacanthodes herrei'' Myers, 1934 * '' Paratriacanthodes retrospinis'' Fowler, 1934 (Sawspine spikefish)

Etymology

''Paratriacanthodes'' prefixes the name of the

type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...

of the Triacanthodidae, '' Triacanthodes'', with ''para'', meaning "near to". This refers to the close relationship between ''Paratriacanthodes'' and ''Triacanthodes''. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

of the type species, ''retrospinis'' means "backward spine" and refers to the backward pointing barb on the first spine in the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

and on the spine in the

pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...

. The specific names of ''P. abei'' and ''P. herrei'' honour the Japanese

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

Tokiharu Abe was a Japanese ichthyologist and a government official of Ministry of Agriculture and Forestry. Career Tokiharu Abe was born in Tabata, Tokyo, emigrated to Taiwan with his family in 1919, graduated from Taipei High School and was admitted to ...

and the American ichthyologist

Albert William Herre Albert William Christian Theodore Herre (September 16, 1868 – January 16, 1962) was an American ichthyologist and lichenologist. Herre was born in 1868 in Toledo, Ohio. He was an alumnus of Stanford University, where he received a Bachelor of ...

respectively.

Characteristics

''Paratriacanthodes'' spikefishes have a flat lower surface to the pelvis, the pelvis tapers to a point to the rear being much broader between the spines of the pelvic fins and at its rear, this feature places the genus in the subfamily Triacanthodinae. These fishes have six spines in the dorsal fin and these descrease in size towards the rear, the smallest sixth spine is, however, still clearly visible. They have a smout which is not as long as the remainder of the ead. The gill slit is short and does extend below the base of the

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

. There is a single row of large conical teeth in each jaw. The largest species in the genus is ''P. retrospinis'' with a maximum published

standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...

of .

Distribution and habitat

''Paratriacanthodes'' spikefishes are found in the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

with ''P. retrospinis'' having a wide distribution being recorded from East Africa, Japan, the Kyushu-Palau Ridge, the South China Sea, Australia and

New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...

. ''P. abei'', however, has only been recorded from the South China Sea and ''P. herrei'' is unknown outside of Philippines waters. These spikefishes are

demersal The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer o ...

and are found at depths between

References

{{Taxonbar, from=Q199222 Triacanthodinae Marine fish genera Taxa named by Henry Weed Fowler