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''Papilio thuraui'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
swallowtail butterfly Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larg ...
from the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Papilio ''Papilio'' is a genus in the swallowtail butterfly family, Papilionidae, as well as the only representative of the tribe Papilionini. The word ''papilio'' is Latin for butterfly. It includes the common yellow swallowtail ('' Papilio machaon'') ...
'' that is found in
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
,
Malawi Malawi, officially the Republic of Malawi, is a landlocked country in Southeastern Africa. It is bordered by Zambia to the west, Tanzania to the north and northeast, and Mozambique to the east, south, and southwest. Malawi spans over and ...
,
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
and the
Republic of the Congo The Republic of the Congo, also known as Congo-Brazzaville, the Congo Republic or simply the Congo (the last ambiguously also referring to the neighbouring Democratic Republic of the Congo), is a country located on the western coast of Central ...
.


Description

Blue median band absent and a complete series of blue submarginal spots in both wings. Both wings above black-brown without median band, but with blue submarginal spots and the forewing also in the middle between the apex of the cell and the distal margin with three small blue spots in cellules 3, 4 and 6; forewing beneath with four large yellowish submarginal spots in cellules 1 b—4. — German East Africa: Ubena
External images


Original description

File:Papilio thuraui Karsch, 1900 Ent. Nachr. 1 (4).jpg , ''Entomologische Nachrichten'' File:Papilio thuraui Karsch, 1900 Ent. Nachr. 1 (2).jpg , File:Papilio thuraui Karsch, 1900 Ent. Nachr. 1 (3).jpg ,


Subspecies

*''P. t. thuraui'' (southern Tanzania, northern Malawi) *''P. t. cyclopis'' Rothschild & Jordan, 1903 . (Malawi, Zambia) *''P. t. occidua'' Storace, 1951 (Republic of the Congo, Malawi, north-eastern Zambia) *''P. t. heathi'' (Hancock, 1984) Hancock, D.L. (1984). The ''Princeps nireus'' group of swallowtails (Lepidoptera: Papilionidae) systematics, phylogeny and biogeography. ''Arnoldia'' Zimbabwe 9(12): 181-215. (Malawi) *''P. t. ngorongoro'' (Hancock, 1984) (northern Tanzania) *''P. t. viphya'' (Hancock, 1984) (Malawi)


Biology

The larvae feed on '' Toddalia asiatica''.


Taxonomy

''Papilio thuraui'' belongs to a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
called the ''nireus''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
with 15 members. The pattern is black with green or blue bands and spots. The butterflies, although called swallowtails, lack tails with the exception of ''Papilio charopus'' and ''Papilio hornimani''. The clade members are: *''
Papilio aristophontes ''Papilio aristophontes'' is a species of butterfly in the family Papilionidae. It is endemic to Comoros. It is listed as a red list species due to its rarity and the threats to its survival. The eggs are laid on the plant '' Toddalia asiatica' ...
'' Oberthür, 1897 *''
Papilio nireus ''Papilio nireus'', the green-banded swallowtail, narrow-banded blue swallowtail, or African blue-banded swallowtail, is a butterfly of the family Papilionidae. It is found in Sub-Saharan Africa. Description The wingspan is in males and in f ...
'' Linnaeus, 1758 *''
Papilio charopus ''Papilio charopus'', the tailed green-banded swallowtail, is a species of swallowtail butterfly from the genus ''Papilio'' that is found in Nigeria, Cameroon, Equatorial Guinea, Republic of the Congo, Uganda, Rwanda, and Burundi. Subspecies *' ...
'' Westwood, 1843 *'' Papilio chitondensis'' de Sousa & Fernandes, 1966 *'' Papilio chrapkowskii'' Suffert, 1904 *''
Papilio chrapkowskoides ''Papilio chrapkowskoides'', the broadly green-banded swallowtail, is a species of swallowtail butterfly from the genus ''Papilio'' that is found in the Republic of the Congo, Uganda, Rwanda, Burundi, Tanzania, Guinea, Sierra Leone, Liberia, Ivo ...
'' Storace, 1952 *'' Papilio desmondi'' van Someren, 1939 *'' Papilio hornimani'' Distant, 1879 *''
Papilio interjectana ''Papilio interjectana'', the Van Someren's green-banded swallowtail, is a species of swallowtail butterfly from the genus ''Papilio'' that is found in Uganda and Kenya.Papilio manlius ''Papilio manlius'' is a species of butterfly in the family Papilionidae. It is endemic to Mauritius. Description The discal band of the forewing in both sexes only composed of three spots (in cellules 3, 4 and in the cell); that of the hindwin ...
'' Fabricius, 1798 *'' Papilio microps'' Storace, 1951 *'' Papilio sosia'' Rothschild & Jordan, 1903 *''Papilio thuraui'' Karsch, 1900 *''
Papilio ufipa ''Papilio ufipa'' is a species of swallowtail butterfly from the genus ''Papilio'' that is found in Tanzania. The habitat consists of riparian and montane forests at altitudes from . Taxonomy ''Papilio ufipa'' belongs to a clade called the ''ni ...
'' Carcasson, 1961 *''
Papilio wilsoni ''Papilio wilsoni'' is a butterfly in the family Papilionidae. It is found in Ethiopia. Taxonomy ''Papilio wilsoni'' belongs to a clade called the ''nireus'' species-group with 15 members. The pattern is black with green bands and spots and ...
'' Rothschild, 1926


Biogeographic realm

This species is located in the
Afrotropical realm The Afrotropical realm is one of the Earth's eight biogeographic realms. It includes Sub-Saharan Africa, the southern Arabian Peninsula, the island of Madagascar, and the islands of the western Indian Ocean. It was formerly known as the Ethiopi ...
.


See also

*
Albertine Rift montane forests The Albertine Rift montane forests ( French: ''Forêts montagnardes du Rift Albertin'') is a tropical moist broadleaf forest ecoregion in east-central Africa. The ecoregion covers the mountains of the northern Albertine Rift, and is home to dis ...
*
Congolian forests The Congolian rainforests ( French: ''Forêts tropicales congolaises'') are a broad belt of lowland tropical moist broadleaf forests which extend across the basin of the Congo River and its tributaries in Central Africa. Description The Congol ...


References

*Carcasson, R.H. (1960). The Swallowtail Butterflies of East Africa (Lepidoptera, Papilionidae). ''Journal of the East Africa Natural History Society'
pdf
Key to East Africa members of the species group, diagnostic and other notes and figures. (Permission to host granted by The East Africa Natural History Society) *


External links


Global Butterfly Information System
Images

Images from
Naturhistorisches Museum Wien The Natural History Museum Vienna () is a large natural history museum located in Vienna, Austria. The NHM Vienna is one of the largest museums and non-university research institutions in Austria and an important center of excellence for all matt ...

External images R C Dening collectionGlobis images
thuraui Butterflies described in 1900 Butterflies of Africa Taxa named by Ferdinand Karsch {{Papilionidae-stub