Papilio Ryukyuensis
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''Papilio bianor'', also known as the common peacock, Chinese peacock black swallowtail emerald or Chinese peacock or bianor's emperor machaon - is a species of
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
in the family
Papilionidae Swallowtail butterflies are large, colorful Butterfly, butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includ ...
, the swallowtails. It is native to Continental Asia. It is the state butterfly of the Indian state of
Uttarakhand Uttarakhand (, ), also known as Uttaranchal ( ; List of renamed places in India, the official name until 2007), is a States and union territories of India, state in North India, northern India. The state is bordered by Himachal Pradesh to the n ...
.


Description

This species is variable in size. Individuals emerged in the spring reach 4 to 8 centimeters wide, while those emerged in the summer can reach 12 centimeters. The forewings are black with dark veining and green scales. The undersides are brown, turning white distally with dark veining. The hindwings are tailed and have ridged edges containing reddish eyespots. The body is black with green scales. The male has black hair on the forewings, which the female lacks.


Description in Seitz

. P. bianor. Geographically as well as seasonally very variable. male: the pilose stripes broad, the posterior three usually united, when they are separated the stripe of the lower median remains broader than the interspace, which separates it from the stripe of the submedian fold; upper surface of the forewing dusted with green or greenblue, this scaling commonly condensed into a band before the narrowly dark distal margin; upper surface of the hindwing anteriorly blue or blue-green and posteriorly dusted with green, commonly with a more thickly scaled large green or blue patch (or a similar band), a red anal spot, besides 0—4 red submarginal spots and at least anteriorly some distinct green submarginal spots. The black basal area of the forewing beneath always extends beyond the base of the upper median, sometimes the whole wing is black; under surface of the hindwing with a complete row of red submarginal spots. The female with less dense metallic scaling; the red spots on the upperside of the hindwing usually more numerous and larger than in the male. The butterfly occurs from North Japan to Tonkin and West China and appears to be fairly common in all its localities, sometimes occurring in profusion. Like the allied species the males congregate on moist sand and visit flowers. The larva is green with 2—8 light oblique bands, on Aurantiaceae. Karl Jordan in Seitz.


Subspecies

There are many subspecies. * ''P. b. okinawensis'' (Yaeyama islands, Okinawa, Japan) * ''P. b. ryukyuensis'' (Okinawa islands, Japan) * ''P. b. amamiensis'' (Amami islands, Kagoshima, Japan) * ''P. b. tokaraensis'' (Tokara islands, Kagoshima, Japan) * ''P. b. hachijonis'' (Hachijo island, Izu islands, Japan) * ''P. b. kotoensis'' Sonan, 1927 (Taiwan) * ''P. b. thrasymedes'' Fruhstorfer 1909 (Taiwan) * ''P. b. polyctor'' Boisduval, 1836 * ''P. b. ganesa'' Doubleday, 1842 (Sikkim, Assam, N.Vietnam, Yunnan) Forewing usually with green discal band, which in specimens from Assam is often only very weakly indicated, but sometimes is broader than in many Sikkim examples. Hindwing with blue patch, which reaches to the 3. radial and is more or less strongly produced distad behind the subcostal and 1. radial. * ''P. b. gladiator'' Fruhstorfer, 1902 unicolorous blue costal area on the upper surface of the hindwing predominates.


Biology

This species can be found in forests and other wooded areas. It can occur in suburban and urban areas if appropriate host plants are available. Food plants include species of
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
, prickly ash, cork trees,
trifoliate orange The trifoliate orange, ''Citrus trifoliata'' ( syn. ''Poncirus trifoliata''), is a member of the family Rutaceae. Whether the trifoliate oranges should be considered to belong to their own genus, ''Poncirus'', or be included in the genus ''Citru ...
,
rue ''Ruta graveolens'', commonly known as rue, common rue or herb-of-grace, is a species of the genus '' Ruta'' grown as an ornamental plant and herb. It is native to the Mediterranean. It is grown throughout the world in gardens, especially for i ...
, and Japanese skimmia. Using the species as a model to investigate the iridescent colour evolution, phylogeography, and the evolution of swallowtail butterflies a chromosome scale genome has been sequenced, the final assembly being 421.52 Mb in size, with 15,375 protein-coding genes and 30 chromosomes (29 autosomes and 1 Z sex chromosome). Phylogenetic analysis of this data indicating that ''P. bianor'' separated from a common ancestor of swallowtails ~23.69–36.04 million years ago.


Taxonomy

It is a member of the species group ''
paris Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, largest city of France. With an estimated population of 2,048,472 residents in January 2025 in an area of more than , Paris is the List of ci ...
''


References


Further reading

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bianor ''Bianor'' is a genus of boreal ecosystem, boreal Salticidae, jumping spiders that can grow to . The robust shiny body and northerly distribution are distinctive. Males can be easily recognized by his swollen forelegs and females have orange le ...
Articles containing video clips Butterflies described in 1777 Butterflies of Indochina Symbols of Uttarakhand Butterflies of Japan Butterflies of Taiwan Taxa named by Pieter Cramer {{Papilionidae-stub