''Papilio helenus'', the red Helen, is a large swallowtail butterfly found in forests of southern

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

and parts of

southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...

Range

''Papilio helenus'' is rarely found in

Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...

, southern and north-east

Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...

Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...

Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...

Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...

Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...

Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...

, Kampuchea,

Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...

, southern

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(including

Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...

Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...

province),

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...

, southern

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...

Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Geography of Taiwan, Taiwan: the Ryukyu Islands are divided into the Satsunan Islands (Ōsumi Islands, Ōsumi, Tokara Islands, Tokara and A ...

, peninsular and eastern

Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...

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Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...

, and

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

(

Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...

Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...

, Bangka,

Kalimantan Kalimantan (; ) is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area, and consists of the provinces of Central Kalimantan, East Kalimantan, North Kalimantan, South Kalimantan, and West Kalimantan. The non-Ind ...

, the

Lesser Sunda Islands The Lesser Sunda Islands (, , ), now known as Nusa Tenggara Islands (, or "Southeast Islands"), are an archipelago in the Indonesian archipelago. Most of the Lesser Sunda Islands are located within the Wallacea region, except for the Bali pro ...

except

Tanimbar The Tanimbar Islands (; ), also called ''Timur Laut'' (literally, "North East"; ), are a group of about 65 islands in the Maluku province of Indonesia. The largest and most central of the islands is Yamdena; others include Selaru to the sout ...

). In India it occurs along the Western Ghats from

Kerala Kerala ( , ) is a States and union territories of India, state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile ...

Gujarat Gujarat () is a States of India, state along the Western India, western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the List of states and union territories ...

, also Palnis and

Shevaroys The Servarayan hills, with the anglicised name Shevaroy Hills, are a towering mountain range (1620 m) near the city of Salem, Tamil Nadu, Salem, in Tamil Nadu state, southern India. The local Tamil language, Tamil name comes from a local deity, ' ...

, in the north from

Mussoorie Mussoorie () is a hill station and a municipal board, in Dehradun city in the Dehradun district of the Indian state of Uttarakhand. It is about from the state capital of Dehradun and north of the national capital of New Delhi. The hil ...

eastwards, to north-east India and onto Myanmar.

Description

Status

''Papilio helenus'' is generally uncommon and slightly threatened ( in certain places) . It is very commonly found Maharashtra, but also rarely in Gujarat and parts of Kerala. Its history can be traced back to Assam and Western Ghats, where it is currently endemic.

Taxonomy

Species group

''Papilio helenus'' is the nominate member of the ''helenus''

species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

. The members of this

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

are: *''Papilio helenus'' Linnaeus, 1758 *'' Papilio iswara'' White, 1842 *'' Papilio iswaroides'' Fruhstorfer, 1898 *'' Papilio nephelus'' Boisduval, 1836 *'' Papilio nubilus'' Staudinger, 1895 *'' Papilio sataspes'' C. & R. Felder, 1865

Subspecies

There are up to thirteen different subspecies, two of which occur in India: * ''P. h. daksha'' Moore - South India. Not rare * ''P. h. helenus'' Linnaeus - Mussoorie to Myanmar. Common and one in Taiwan: * ''P. h. fortunius'' Fruhstorfer, 1908

Life cycle

This butterfly flies throughout the year in southern India.

Eggs

The

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...

is pale

apricot An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus ''Prunus''. Usually an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are also ...

-yellow in colour when freshly deposited, spherical in shape and has a slightly roughened exterior which looks like the skin of an orange when seen through a

microscope A microscope () is a laboratory equipment, laboratory instrument used to examine objects that are too small to be seen by the naked eye. Microscopy is the science of investigating small objects and structures using a microscope. Microscopic ...

. The diameter of an egg is 1.2 mm. The eggs are deposited singly on the tips of very young

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

and shoots in shady parts of thick jungle. Before hatching, the eggs appear to be marked by chocolate coloured lines and flecks. The egg hatches in 4 to 7 days.

Caterpillar

The freshly emerged larva is about 3 mm long. Throughout the caterpillar stage, if it is agitated it can evert a yellow-to-red osmeterium from the first segment, just behind the head. Each of the other segments bears, on the back, a pair of tufts of stiff hairs, each tuft arising from a small, yellowish conical process. The overall colour is brown, but there is a whitish

saddle A saddle is a supportive structure for a rider of an animal, fastened to an animal's back by a girth. The most common type is equestrian. However, specialized saddles have been created for oxen, camels and other animals. It is not know ...

-like patch about the middle and the tail segments are also whitish in colour. Like other ''Papilio'' species the larva can evert a two-pronged horn-like osmeterium when it is irritated. The osmeterium secretes an repugnatorial, unpleasant-smelling liquid which is believed to repel

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

s and

parasites Parasitism is a close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The en ...

. After the first moult the

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

has the appearance of a shiny bird dropping. The larva is grass green in colour, mottled black and white and smoky grey. The osmeterium, when everted, is generally yellow to red. While inactive, mainly during daylight hours, the young larva lies along the midrib of the underside of the leaf. Later on, when it is largely fully grown, it is greener and lies on the centre of the upperside of the leaf, on a stem or a twig. The fifth

instar An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to ...

larva is about 5 cm long.

References

* * * * * *

External links

* {{Taxonbar, from=Q669913 helenus Butterflies of Malaysia Butterflies described in 1758 Butterflies of Indochina Taxa named by Carl Linnaeus Butterflies of Sri Lanka Butterflies of Taiwan Butterflies of Japan Butterflies of Java Butterflies of Borneo Lepidoptera of the Philippines Lepidoptera of Sumatra