Paperbark Flycatcher
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The paperbark flycatcher (''Myiagra nana''), also known as the little restless flycatcher, is a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird in the family
Monarchidae The monarchs or monarch flycatchers (family Monarchidae) comprise a family of over 100 passerine birds which includes shrikebills, paradise flycatchers, and Grallina, magpie-larks. Monarchids are small insectivore, insectivorous songbirds with l ...
. It occurs in tropical woodland and riverine habitats of northern Australia and southern New Guinea. Previously, some authorities lumped the paperbark flycatcher as a distinctive
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
restless flycatcher The restless flycatcher (''Myiagra inquieta'') or restless myiagra is a passerine bird in the family Monarchidae; it is also known as the razor grinder or scissors grinder because of its distinctive call. It is a native of eastern and southern A ...
(''Myiagra inquieta'') of southern and eastern Australia, with which it forms a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
.


Taxonomy and systematics

Originally described as ''Seisura nana'' by
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist who published monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould (illustrator), Elizabeth Gould, and several other artists, includ ...
in 1870, the paperbark flycatcher was long treated as a subspecies of the
restless flycatcher The restless flycatcher (''Myiagra inquieta'') or restless myiagra is a passerine bird in the family Monarchidae; it is also known as the razor grinder or scissors grinder because of its distinctive call. It is a native of eastern and southern A ...
(''M. inquieta''), until 1999 when Schodde and Mason identified the distinctness and lack of intergrading in closely situated populations of the two parapatric
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
.


Description

The paperbark flycatcher is broadly similar to the restless flycatcher, with entirely black upperparts from the crown and sides of the head, in contrast with entirely white underparts from the throat to the vent. It is a smaller bird, at only two-thirds the weight of its southern relative, and has a proportionately shorter and broader bill, with longer and stouter rictal bristles. There is no overlap in size between the species. The back and the crown of ''nana'' are the same glossy black, while ''inquieta'' has a slightly paler, slate-grey, back. There are also differences in the vocalisations: ''nana'' almost never use the distinctive "scissors-grinding" call of ''inquieta''.


Distribution and habitat

The flycatcher is found in northern Australia, from the
Kimberley region The Kimberley is the northernmost of the nine regions of Western Australia. It is bordered on the west by the Indian Ocean, on the north by the Timor Sea, on the south by the Great Sandy Desert, Great Sandy and Tanami Desert, Tanami deserts in t ...
of Western Australia, across the
Top End The Top End of Australia's Northern Territory is a geographical region encompassing the northernmost section of the Northern Territory, which aside from the Cape York Peninsula is the northernmost part of the Australian continent. It covers a ...
of the
Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...
, to the
Gulf Country The Gulf Country or North West Queensland is the region of woodland and savanna grassland surrounding the Gulf of Carpentaria in north western Queensland and eastern Northern Territory on the north coast of Australia. The region is also ca ...
and south-western
Cape York Peninsula The Cape York Peninsula is a peninsula located in Far North Queensland, Australia. It is the largest wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth's last wild places. Mexico City: Agrupación Sierra Madre, ...
of north-west
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
, with a population on
Saibai Island Saibai Island, commonly called Saibai (), is an island of the Torres Strait Islands archipelago, located in the Torres Strait of Queensland, Australia. The island is situated north of the Australian mainland and south of the island of New Guine ...
in
Torres Strait The Torres Strait (), also known as Zenadh Kes ( Kalaw Lagaw Ya#Phonology 2, ˆzen̪ad̪ kes, is a strait between Australia and the Melanesian island of New Guinea. It is wide at its narrowest extent. To the south is Cape York Peninsula, ...
. It is also found in southern
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
from
Merauke Merauke is a large town (''kelurahan'') and an administrative district (''distrik'') in Merauke Regency of South Papua Province, Indonesia. It is also the administrative centre of Merauke Regency, and is considered to be the easternmost city in I ...
eastwards to the Bensbach River in the Middle Fly District. The paperbark flycatcher does not overlap in distribution with the restless; although their ranges abut in northern Queensland there are no signs of intergradation between the two species, and in the area where the ranges meet most of the restless flycatchers are non-breeding migrants. In Australia the paperbark flycatcher inhabits tropical
eucalypt Eucalypt is any woody plant with Capsule (fruit), capsule fruiting bodies belonging to one of seven closely related genera (of the tribe Eucalypteae) found across Australia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', ''Stockwellia'', ''Allosyn ...
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s, paperbark woodlands and dry riverine woodlands. In New Guinea, where it is locally common along the lower reaches of the Bensbach River, it frequents scrub, partly submerged trees, and sedgeland bordering rivers on
floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river. Floodplains stretch from the banks of a river channel to the base of the enclosing valley, and experience flooding during periods of high Discharge (hydrolog ...
s and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
.


Behaviour

A conspicuous bird, the paperbark flycatcher is usually seen in pairs or singly. It sweeps its tail restlessly from side to side and often hovers near the ground when searching for food.


Breeding

Nesting habits in New Guinea are undescribed. In northern Australia breeding takes place from November to January. The nest is a neat cup built of bark shreds and grass stems in the fork of a dead shrub near water. One whitish egg, spotted reddish-brown and grey, is laid.


References


Notes


Sources

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External links


Egg of paperbark flycatcher
{{Taxonbar, from=Q1923015 paperbark flycatcher Birds of the Northern Territory Birds of Queensland Birds of Western Australia Birds of southern New Guinea paperbark flycatcher