''Panna'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the family

Sciaenidae Sciaenidae is a family (biology), family of ray-finned fishes belonging to the Order (biology), order Acanthuriformes. They are commonly called drums or croakers in reference to the repetitive throbbing or drumming sounds they make. The family co ...

, the drums and croakers. These fishes are found in southern and southeast Asia.

Taxonomy

''Panna'' was first proposed as a

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

in 1969 by the Indian

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

R. S. Lal Mohan with ''Otolithus microdon'' designated as its

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, as well as being the only species. ''O. microdon'' was described in 1849 by

Pieter Bleeker Pieter Bleeker (10 July 1819 – 24 January 1878) was a Dutch medical doctor, Ichthyology, ichthyologist, and Herpetology, herpetologist. He was famous for the ''Atlas Ichthyologique des Indes Orientales Néêrlandaises'', his monumental work on ...

, with its type locality given as Java. The genus ''Pachyurus'' is included in the subfamily Cynoscioninae by some workers, but the 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' does not recognise subfamilies within the Sciaenidae, which it places in the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Acanthuriformes Acanthuriformes is a large, diverse order of mostly marine ray-finned fishes, part of the Percomorpha clade. In the past, members of this clade were placed in the suborders Acanthuroidea and Percoidea of the order Perciformes, but this treatment ...

Etymology

''Panna'' is a name that Lal Mohan did not explain but is thought to be a local name of the type species, ''P. macrodon'' in India.

Species

There are currently 3 recognized species in this genus: * '' Panna heterolepis'' Trewavas, 1977 (Hooghly croaker) * ''

Panna microdon Panna can refer to: * Paññā is Pali for "wisdom"; the Sinhala version is ''Prajñāva'' Food * Aam panna, an Indian drink made from mangoes * Panna cotta ("cooked cream"), an Italian dessert * Panna (water), an Italian bottled water Place ...

'' ( Bleeker, 1849) (Panna croaker) * '' Panna perarmatus'' ( Chabanaud, 1926) (Armour croaker)

Characteristics

''Panna'' croakers have a

swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...

characterised by having a pair of tubular appendages starting at the head end which divides almost at its root into a simple rear part and either a simple or branched part at the head end, entering the head through the

septum transversum The septum transversum is a thick mass of cranial mesenchyme, formed in the embryo, that gives rise to parts of the thoracic diaphragm and the ventral mesentery of the foregut in the developed human being and other mammals. Origins The septum t ...

. The maximum published

standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...

s of ''Panna'' croakers varies from in ''P. heterolepis'' and in ''P. perarmatus''.

Distribution

''Panna'' croakers are found in the

Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the s ...

with ''P. heterolepis'' being found off India, Sri Lanka, Bangladesh and Myanmar; ''P. microdon'' is found in eastern Malaysia in

Perak Perak (; Perak Malay: ''Peghok'') is a States and federal territories of Malaysia, state of Malaysia on the west coast of the Malay Peninsula. Perak has land borders with the Malaysian states of Kedah to the north, Penang to the northwest, Kel ...

, the

Gulf of Thailand The Gulf of Thailand (), historically known as the Gulf of Siam (), is a shallow inlet adjacent to the southwestern South China Sea, bounded between the southwestern shores of the Indochinese Peninsula and the northern half of the Malay Peninsula. ...

, Vietnam,

Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...

, eastern

Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...

and

Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...

; and ''P. perarmatus'' is found off Vietnam, in the Gulf of Thailand and Borneo.

References

{{Taxonbar, from=Q11846406 Sciaenidae