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''Pangasius'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of medium-large to very large
shark catfish The shark catfishes form the family Pangasiidae. They are found in fresh and brackish waters across southern Asia, from Pakistan to Borneo. Among the 30-odd members of this family is the plant-eating, endangered Mekong giant catfish ''Pangasia ...
es native to fresh water in
South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. The term "pangasius" is sometimes used to specifically refer to the commercially important
basa fish Basa (''Pangasius bocourti'') is a species of catfish in the family Pangasiidae. Basa are native to the Mekong and Chao Phraya basins in Mainland Southeast Asia. These fish are important food fish with an international market. They are often ...
, ''P. bocourti''.


Taxonomy

In 1993, ''Pangasius'' was one of two extant genera (along with '' Helicophagus'') in the family Pangasiidae. At this point, it was split into four subgenera. ''Pangasius'' (''Pangasianodon'') included '' P. gigas'' and '' P. hypophthalmus'' and was diagnosed by the absence of mandibular barbels, the absence of teeth in adults and the presence of a single-lobed
swimbladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...
. ''Pangasius'' (''Pteropangasius'') included ''P. micronema'' and ''P. pleurotaenia'' and was diagnosed by four lobes in the swimbladder and with multiple segments in the last lobe. ''Pangasius'' (''Neopangasius'') included ''P. nieuwenhuisii'', ''P. humeralis'', ''P. lithostoma'', ''P. kinabatanganensis'', diagnosed by
palatal The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...
teeth arranged in a single large patch and high
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characterist ...
l counts. ''Pangasius'' (''Pangasius'') was the final subgenus and had no diagnostic features, containing the remaining species. These subgeneric classifications were confirmed in 2000 except for ''Neopangasius'', found to be
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
and to be part of ''Pangasius'' (''Pangasius''), thus leaving three subgenera. Since then, the subgenera have been variably recognized as separate. ''P. gigas'' and ''P. hypophthalmus'' have been classified in the genus ''
Pangasianodon ''Pangasianodon'' is a genus of large to very large shark catfishes native to the Mekong and Chao Phraya Rivers in Southeast Asia and adjacent China. Species Two species in this genus are recognized: * '' Pangasianodon gigas'' Chevey, 1931 (M ...
'', and ''P. micronemus'' and ''P. pleurotaenia'' in the genus ''
Pseudolais ''Pseudolais'' is a genus of shark catfishes native to Southeast Asia. Species There are currently two recognized species in this genus: * ''Pseudolais micronemus'' ( Bleeker, 1846) (Shortbarbel pangasius) * ''Pseudolais pleurotaenia'' (Sauvage ...
'' (with ''Pteropangasius'' as a junior synonym). In 2011, ''Pangasius'' was sixth in the
National Fisheries Institute The National Fisheries Institute (NFI) is the United States industry trade group representing the seafood industry. It is a member of the International Coalition of Fisheries Associations (ICFA). Its member companies consist of all levels of busi ...
’s "Top 10" list of the most consumed seafood in the United States. The Top 10 is based on tonnage of fish sold. According to the NFI, this mild-flavored white-fleshed fish is farmed in Asia, and is being used increasingly in food service. It is finding its way onto restaurant menus and into stores, as well, where one may see it called basa, tra, or swai. They are either called Panga, Pangas or Pangasius, In
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
and
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, Pangasius are called Ikan Patin , while Malaysian Chinese call Pangasius 巴丁鱼. Some species like Pangasius Nasutus , Pangasius Djambal and Pangasius Sanitwongsei are expensive food fish in
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
, Pangasius Sanitwongsei was also a common fish in aquarium trade and
sport fishing Recreational fishing, also called sport fishing or game fishing, is fishing for leisure, exercise or competition. It can be contrasted with commercial fishing, which is professional fishing for profit; or subsistence fishing, which is fishin ...
.


Species

Currently, 22 recognized species are in this genus: * ''
Pangasius bocourti Basa (''Pangasius bocourti'') is a species of catfish in the family Pangasiidae. Basa are native to the Mekong and Chao Phraya basins in Mainland Southeast Asia. These fish are important food fish with an international market. They are often ...
''
Sauvage Sauvage, French for "savage" may refer to: ; as a surname: * Catherine Sauvage (1929–1998), a French singer and actress * Cyrille Sauvage (born 1973), a French racing driver * Frédéric Sauvage (1786–1857), a French boat builder who carried ...
, 1880
(basa fish) * ''
Pangasius conchophilus ''Pangasius conchophilus'' is a species of shark catfish.Roberts, Tyson R., and Chavalit Vidthayanon. "Systematic revision of the Asian catfish family Pangasiidae, with biological observations and descriptions of three new species." Proceedings ...
''
Roberts Roberts may refer to: People * Roberts (given name), a Latvian masculine given name * Roberts (surname), a popular surname, especially among the Welsh Places * Roberts (crater), a lunar impact crater on the far side of the Moon ;United Stat ...
& Vidthayanon, 1991
(snail eating pangasius) * ''
Pangasius djambal ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. ...
''
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Pangasius elongatus'' Pouyaud, Gustiano & Teugels, 2002 (elongated pangasius) * '' Pangasius humeralis'' Roberts, 1989 * ''
Pangasius kinabatanganensis ''Pangasius kinabatanganensis'' is a species of shark catfish.Roberts, Tyson R., and Chavalit Vidthayanon. "Systematic revision of the Asian catfish family Pangasiidae, with biological observations and descriptions of three new species." Proceed ...
'' Roberts & Vidthayanon, 1991 (kinabatang pangasius) * ''
Pangasius krempfi ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South Asia, South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. boco ...
''
Fang A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fan ...
& Chaux, 1949
* '' Pangasius kunyit'' Pouyaud, Teugels &
Legendre Legendre, LeGendre or Le Gendre is a French surname. It may refer to: * Adrien-Marie Legendre (1752–1833), French mathematician ** Associated Legendre polynomials ** Legendre's equation ** Legendre polynomials ** Legendre symbol ** Legendre tran ...
, 1999
* ''
Pangasius larnaudii ''Pangasius larnaudii'', the black ear catfish is a species of freshwater shark catfish endemic to Mekong The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's twelfth longest river an ...
'' Bocourt, 1866 (spot pangasius) * ''
Pangasius lithostoma ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. T ...
''
Roberts Roberts may refer to: People * Roberts (given name), a Latvian masculine given name * Roberts (surname), a popular surname, especially among the Welsh Places * Roberts (crater), a lunar impact crater on the far side of the Moon ;United Stat ...
, 1989
* ''
Pangasius macronema ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. T ...
'' Bleeker, 1851 * ''
Pangasius mahakamensis ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. T ...
'' Pouyaud, Gustiano & Teugels, 2002 * ''
Pangasius mekongensis ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. ...
'' Gustiano, Teugels & Pouyaud, 2003 (mekong pangasius) * '' Pangasius myanmar'' Roberts & Vidthayanon, 1991 (myanmar pangasius) * ''
Pangasius nasutus ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. ...
'' (
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Pangasius nieuwenhuisii ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. Tax ...
'' (
Popta Canna Maria Louise Popta (31 May 1860 – 13 June 1929)L.B. Holthuis, Biography i1820-1958, Rijksmuseum van Natuurlijke Historie page 82 (in Dutch). was a Dutch biologist. Born in Breda, Popta was one of the first women to enrol as a student at ...
, 1904)
* '' Pangasius pangasius'' ( Hamilton, 1822) (yellowtail catfish) * ''
Pangasius polyuranodon ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. ...
'' Bleeker, 1852 * ''
Pangasius rheophilus ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. ...
'' Pouyaud & Teugels, 2000 * ''
Pangasius sabahensis ''Pangasius'' is a genus of medium-large to very large shark catfishes native to fresh water in South and Southeast Asia. The term "pangasius" is sometimes used to specifically refer to the commercially important basa fish, ''P. bocourti''. T ...
'' Gustiano, Teugels & Pouyaud, 2003 * ''
Pangasius sanitwongsei The giant pangasius, paroon shark, pangasid-catfish or Chao Phraya giant catfish (''Pangasius sanitwongsei'') is a species of freshwater fish in the shark catfish family (Pangasiidae) of order Siluriformes, found in the Chao Phraya and Mekong ...
''
Smith Smith may refer to: People * Metalsmith, or simply smith, a craftsman fashioning tools or works of art out of various metals * Smith (given name) * Smith (surname), a family name originating in England, Scotland and Ireland ** List of people w ...
, 1931
(giant pangasius) * '' Pangasius silasi'' Dwivedi et al., 2017


Fossil record

The single known fossil species of this genus, '' P. indicus'', is reported from the
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
period of Sipang, Sumatra, either from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
or the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
.


References

{{Taxonbar, from=Q312329 Pangasiidae Paleogene genus first appearances Catfish genera Taxa named by Achille Valenciennes Freshwater fish genera Extant Paleogene first appearances