Palmnut Vulture
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The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
in the family
Accipitridae The Accipitridae () is one of the four families within the order Accipitriformes, and is a family of small to large birds of prey with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects ...
(which also includes many other diurnal
raptor Raptor(s) or RAPTOR may refer to: Animals The word "raptor" refers to several groups of avian and non-avian dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunt ...
s such as
kite A kite is a tethered heavier than air flight, heavier-than-air craft with wing surfaces that react against the air to create Lift (force), lift and Drag (physics), drag forces. A kite consists of wings, tethers and anchors. Kites often have ...
s,
buzzard Buzzard is the common name of several species of birds of prey. ''Buteo'' species * Archer's buzzard (''Buteo archeri'') * Augur buzzard (''Buteo augur'') * Broad-winged hawk (''Buteo platypterus'') * Common buzzard (''Buteo buteo'') * Easte ...
s and harriers, vultures, and eagles). It is the only member of the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Gypohierax''. This bird is an
Old World vulture Old World vultures are vultures that are found in the Old World, i.e. the continents of Europe, Asia and Africa, and which belong to the family Accipitridae, which also includes eagles, buzzards, kites, and hawks. Old World vultures are not ...
(only distantly related to the
New World vulture Cathartidae, known commonly as New World vultures or condors, are a family (biology), family of birds of prey consisting of seven extant species in five genus, genera. It includes five extant vultures and two extant condors found in the Americas. ...
s, which are in a separate family, the
Cathartidae Cathartidae, known commonly as New World vultures or condors, are a family of birds of prey consisting of seven extant species in five genera. It includes five extant vultures and two extant condors found in the Americas. They are known as "New W ...
). It breeds in forest and savannah across
sub-Saharan Africa Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
, usually near water, its range coinciding with that of the
oil An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) and lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturate ...
and
Raffia palm Raffia palms are members of the genus ''Raphia''. The Malagasy name is derived from ' "to squeeze juice". The genus contains about twenty species of palms native to tropical regions of Africa, and especially Madagascar, with one species ('' R ...
s. It is quite approachable, like many African vultures, and can be seen near habitation, even on large hotel lawns in the tourist areas of countries such as
the Gambia The Gambia, officially the Republic of The Gambia, is a country in West Africa. Geographically, The Gambia is the List of African countries by area, smallest country in continental Africa; it is surrounded by Senegal on all sides except for ...
.


Taxonomy

The palm-nut vulture was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
in 1788 by the German naturalist
Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
's ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. He placed it with the eagles, hawks and falcons in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Falco'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Falco angolensis''. Gmelin based his description on the "Angola vulture" that had been described in 1781 by the English ornithologist John Latham in his multi-volume work ''A General Synopsis of Birds''. The
Leverian Collection The Leverian collection was a natural history and ethnographic collection assembled by Ashton Lever. It was noted for the content it acquired from the voyages of Captain James Cook. For three decades it was displayed in London, being broken up ...
in London included two specimens that had been collected in Angola. The palm-nut vulture is now placed in the genus ''Gypohierax'' that was introduced for the species in 1836 by the German naturalist
Eduard Rüppell Wilhelm Peter Eduard Simon Rüppell, also spelled Rueppell (20 November 1794 – 10 December 1884) was a German Natural history, naturalist and List of explorers, explorer, best known for his collections and descriptions of plants and animals from ...
. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''gups'' meaning "vulture" with ''hierax'' meaning "hawk". The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

This is a nearly unmistakable bird as an adult. At in weight, in length and with a wingspan of , this is the smallest
Old World vulture Old World vultures are vultures that are found in the Old World, i.e. the continents of Europe, Asia and Africa, and which belong to the family Accipitridae, which also includes eagles, buzzards, kites, and hawks. Old World vultures are not ...
. Its plumage is all white except for black areas in its wings and tail. It has a red patch around each eye. The juvenile, which takes 3–4 years to mature, is brown with yellow eye-patches. In flight, this species resembles an eagle more than a typical vulture, and it can sustain flapping flight, so it does not depend on thermals. With its extensive white plumage, and black wing- and tail-feathers, the adult palm-nut vulture can be crudely mistaken for both the African fish-eagle and the
Egyptian vulture The Egyptian vulture (''Neophron percnopterus''), also called the white scavenger vulture or pharaoh's chicken, is a small Old World vulture in the monotypic genus ''Neophron''. It is widely distributed from the Iberian Peninsula, North Africa ...
, but clearly lacks the chestnut body of the former and the white tail of the latter. The sexes are identical in appearance, with the female being the same size as the male. Juveniles on the other hand are predominately brown with partially black wings and take a lengthy three to four years to make the transition into the adult plumage.


Distribution and habitat

Palm-nut vultures are found throughout most of the coastal areas of the African continent from The Gambia to Kenya and as far South as South Africa. The total African population is estimated to be 80,000 pairs. There are approximately 40 birds in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. The only Southern African subregions to have resident breeding pairs of palm-nut vultures are South Africa and Mozambique. The breeding distribution of the palm-nut vulture during the 1970s census period was centred on the
Raffia palm Raffia palms are members of the genus ''Raphia''. The Malagasy name is derived from ' "to squeeze juice". The genus contains about twenty species of palms native to tropical regions of Africa, and especially Madagascar, with one species ('' R ...
groves of the
Kosi Bay Kosi Bay is a series of four interlinked lakes in the Maputaland area of KwaZulu-Natal, South Africa. Ecology The lakes form part of the iSimangaliso Wetland Park, a UNESCO World Heritage Site. The closest town is Manguzi, some away from it. ...
system and
Mtunzini Mtunzini ( Zulu: ''eMthunzini'', from ''umthunzi'' meaning "a place in the shade") is a small coastal town that is situated almost exactly halfway along KwaZulu-Natal's coastline in South Africa approximately 140 km north of Durban. In 2011, t ...
. Its distribution is linked to the presence of the Raffia palm ''
Raphia australis ''Raphia australis'', the giant palm, Kosi Palm, or rafia, is a species of raffia palm in the family Arecaceae. It is found around Kosi Bay in southern Mozambique and northeastern KwaZulu Natal in South Africa. It is threatened by habitat loss c ...
'' at all permanently occupied sites, and the existence of this species at Mtunzini is entirely due to the artificial cultivation of Raffia palms. Currently there are 7 known nesting sites in South Africa and a total of 40 individual birds. As the name suggests, the distribution of the palm-nut vulture closely tracks that of oil (''
Elaeis guineensis ''Elaeis guineensis'' is a species of Arecaceae, palm commonly just called oil palm but also sometimes African oil palm or macaw-fat. The first Western world, Western person to describe it and bring back seeds was the French naturalist Michel Ad ...
'') or raffia ('' Raphia'' sp.) palms. Consequently, it is most common in coastal forests and mangrove swamps below , but also occurs in wet savannas.


Behaviour and ecology


Food and feeding

Unusual for birds of prey, the palm-nut vulture feeds mainly on the fleshy fruit-husks of the oil palm and on the palm-fruits of Raffia palms, as well as wild dates,
orange Orange most often refers to: *Orange (fruit), the fruit of the tree species '' Citrus'' × ''sinensis'' ** Orange blossom, its fragrant flower ** Orange juice *Orange (colour), the color of an orange fruit, occurs between red and yellow in the vi ...
s, other
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
s, some
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
s and
acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s.Ferguson-Lees, J. & Christie, D.A. & Franklin, K. & Mead, D. & Burton, P.. (2001). Raptors of the world. Helm Identification Guides. These fruits make up over 60% of the adult bird's diet and over 90% of the juvenile bird's diet. It has also been recorded to feed on crabs (both freshwater and marine),
molluscs Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
, frogs,
tadpole A tadpole or polliwog (also spelled pollywog) is the Larva, larval stage in the biological life cycle of an amphibian. Most tadpoles are fully Aquatic animal, aquatic, though some species of amphibians have tadpoles that are terrestrial animal, ...
s, fish,
dung beetle Dung beetles are beetles that feed on feces. All species of dung beetle belong to the superfamily Scarabaeoidea, most of them to the subfamilies Scarabaeinae and Aphodiinae of the family Scarabaeidae (scarab beetles). As most species of Scara ...
s, termites, alate
ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s, alate termites,
locust Locusts (derived from the Latin ''locusta'', locust or lobster) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances they b ...
s, small mammals,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s and their nestlings, snakes, other
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s, even reptiles’ eggs and hatchlings, and it has been known to occasionally attack domestic poultry and feed on
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
.


Breeding

Breeding pairs construct large stick nests high up in tall trees and will often exhibit a strong attachment to the nest site. They may stay at the nesting site for an entire year. Where Raphia Palms are present, breeding pairs will build a nest at the base of the palm fronds. At the beginning of the breeding season, pairs soar together in an aerial display of rolling and diving, much more acrobatic than most vultures. During each breeding cycle, a single, white and brown egg is laid, which is incubated by both sexes, over a period of four to six weeks. Normally around 85 to 90 days after hatching, the young brown chicks will fledge.


Conservation status

This species is widespread throughout much of Africa, overall fairly common and with a stable population. It is rarer and much more localized in South Africa, although not considered to be under any form of immediate threat in that country. That being said, the threats to this species in South Africa are not well understood. The low population size makes the species vulnerable to stochastic events. In Mozambique Parker (1999) observed that cutting down of coastal forest was threatening breeding habitat. The main threat to this species in South Africa is habitat loss. Open cast sand dune mining and urban expansion have reduced suitable habitats. The cultivation of Raphia palms for its ornamental value is currently increasing providing food and nesting sites. There is also a large portion of its habitat protected by the
Isimangaliso Wetland Park iSimangaliso Wetland Park is situated on the east coast of KwaZulu-Natal, South Africa, about north of Durban by road. It is South Africa's third-largest protected area, spanning of coastline, from the Mozambique, Mozambican border in the n ...
. There are no current species specific conservation initiatives as this species is the only vulture species in South Africa where the population size is increasing. The importance of maintaining this is that the South African satellite population in producing dispersing birds that are contributing to maintaining the Mozambique population (and hence the larger metapopulation). File:Palmiste africain.jpg, Head of palm-nut vulture Laika ac Selous (9859653096).jpg, Immature bird in
Selous Game Reserve The Selous Game Reserve, now renamed as Nyerere National Park (in-part), is a protected nature reserve and wilderness area in southern Tanzania, East Africa. It covers a total area of , with additional buffer zones, as well. It was designated a ...
, Tanzania Palm nut vulture (Gypohierax angolensis) in flight, Semliki Wildlife Reserve.jpg, Adult in flight in
Semliki Wildlife Reserve The Semliki Wildlife Reserve is a conservation protected area in the Western Region of Uganda with headquarters at Karugutu in Ntoroko District. It is the oldest wildlife reserve. Geography Semliki Wildlife Reserve is located in Kabarole an ...
, Uganda Palm nut vulture (Gypohierax angolensis) on nest.jpg, Adult on nest at
Kazinga Channel The Kazinga Channel in Uganda is a wide, long natural channel that links Lake Edward and Lake George (Uganda), Lake George, which are part of the African Great Lakes system. The channel is a dominant feature of Queen Elizabeth National Park. ...
, Uganda


References


Further reading

*


External links

* Palmnut Vulture
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q732523
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the ...
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the ...
Birds of the Gulf of Guinea Birds of prey of Sub-Saharan Africa
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the ...
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the ...