Paleotis
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''Palaeotis'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
paleognath Palaeognathae (; ) is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Pala ...
birds from the middle
Eocene epoch The Eocene ( ) is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''Ēṓs'', 'Dawn') and ...
of central Europe. One species is known, ''Palaeotis weigelti''. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen is a fossil
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
and
phalanx The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
. Lambrect (1928) described it as an extinct
bustard Bustards, including floricans and korhaans, are large, terrestrial birds living mainly in dry grassland areas and in steppe regions. They range in length from . They make up the family Otididae (, formerly known as Otidae). Bustards are ...
(genus ''Otis''), and gave it its consequent name (''Palaeotis'' means ''ancient bustard'').Lambrecht, K. (1928) ''Palaeotis weigelti n. g. sp., eine fossil trappe aus der mitteleozanen Braunkohle des Geiseltales''. Jahrbuch hallesch. Verband., Halle, n.s., 7:11. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that ''Palaeotis'' is a palaeognath and assigned it to the same
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
as
ostrich Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large parts of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa. They are the heaviest and largest living birds, w ...
es; the
Struthioniformes Struthioniformes is an order of birds with only a single extant family, Struthionidae, containing the ostriches. Several other extinct families are known, spanning across the Northern Hemisphere, from the Early Eocene to the early Pliocene, incl ...
.Houde, Peter. Haubold, Hartmut. (1987) Palaeovertebrata, 17 (2): 27 - 46, 6 figures. 20 June 1987. In 2021, it was considered a member of the family Paleotididae alongside ''
Galligeranoides ''Galligeranoides'' is an extinct genus of geranoidid that lived during the Ypresian. Distribution ''Galligeranoides boriensis'' is known from the site of La Borie in France France, officially the French Republic, is a country located p ...
'' from the Early Eocene of France, which were found to be basal members of the Struthioniformes. In the 1930s a nearly complete fossil with catalog number GM 4362 was assigned to ''Palaeotis'', probably by Lambrecht. Houde and Haubold found three additional specimens in the collection of the Geiseltalmuseum, Martin-Luther University, Halle/S., Germany. One of those three is the holotype specimen of ''Paleogrus geiseltalensis'' (=Ornithocnemus geiseltalensis, Lambrecht 1935). Houde and Haubold also requested permission to prepare a fossil cataloged as HLMD Me 7530 at the Hesseches Landesmuseum in Darmstadt, Germany. HLMD Me 7530 was collected from the famous
Messel shales The Messel Formation is a geologic formation in Hesse, central Germany, dating back to the Eocene epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlies crystalline Variscan basement and it ...
. When it was prepared, the two ornithologists assigned it to ''Palaeotis'' as well. Other scientists are less convinced that ''Palaeotis'' is a struthioniform, placing it instead as a more basal
ratite Ratites () are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal ...
. It may be related to the mysterious '' Remiornis'', a putative ratite known from the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
. Various other ratite remains also occur in the European Paleogene and early Miocene, and these may represent various independent lineages, leading to further confusion.


Biology

''Palaeotis'' is suggested to be sexually dimorphic, as some specimens being consistently smaller than others. While the forelimb is incomplete, the spindley humerus indicates that it had relatively large wings unlike modern
ostriches Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large parts of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa. They are the heaviest and largest living birds, w ...
and
rheas The rhea ( ), also known as the ñandu ( ) or South American ostrich, is a South American ratite (flightless bird without a keel (bird anatomy), keel on the sternum bone) of the order Rheiformes. They are distantly related to the two African Ost ...
. The beak was slender, more similar to that of lithornithids, and probably had similar carnivorous habits. ''Paleotis'' is considered to be
flightless Flightless birds are birds that cannot fly, as they have, through evolution, lost the ability to. There are over 60 extant species, including the well-known ratites ( ostriches, emus, cassowaries, rheas, and kiwis) and penguins. The smal ...
.


References

Eocene birds of Europe Struthioniformes {{paleo-bird-stub