Cave of Altamira and Paleolithic Cave Art of Northern Spain-110113

The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός '' palaios'', "old" and λίθος ''

lithos Lithos is a glyphic sans-serif typeface designed by Carol Twombly in 1989 for Adobe Systems. Lithos is inspired by the unadorned, geometric letterforms of the engravings found on Ancient Greek public buildings. The typeface consists of only ca ...

'', "stone"), is a period in human

prehistory Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use ...

that is distinguished by the original development of

stone tool A stone tool is, in the most general sense, any tool made either partially or entirely out of stone. Although stone tool-dependent societies and cultures still exist today, most stone tools are associated with prehistoric (particularly Stone Ag ...

s, and which represents almost the entire period of human prehistoric technology. It extends from the earliest known use of stone tools by hominins, 3.3 million years ago, to the end of the

Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...

, 11,650

cal Cal or CAL may refer to: Arts and entertainment * ''Cal'' (novel), a 1983 novel by Bernard MacLaverty * "Cal" (short story), a science fiction short story by Isaac Asimov * ''Cal'' (1984 film), an Irish drama starring John Lynch and Helen Mir ...

BP. The Paleolithic Age in Europe preceded the Mesolithic Age, although the date of the transition varies geographically by several thousand years. During the Paleolithic Age, hominins grouped together in small societies such as bands and subsisted by gathering plants, fishing, and hunting or scavenging wild animals. The Paleolithic Age is characterized by the use of knapped

s, although at the time humans also used wood and bone tools. Other organic commodities were adapted for use as tools, including

leather Leather is a strong, flexible and durable material obtained from the tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffalo, pigs and ho ...

and vegetable

fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorpora ...

s; however, due to rapid decomposition, these have not survived to any great degree. About 50,000 years ago, a marked increase in the diversity of artifacts occurred. In Africa, bone artifacts and the first art appear in the archaeological record. The first evidence of human

fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques ...

is also noted, from artifacts in places such as Blombos cave in

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...

. Archaeologists classify artifacts of the last 50,000 years into many different categories, such as

projectile point In North American archaeological terminology, a projectile point is an object that was hafted to a weapon that was capable of being thrown or projected, such as a javelin, dart, or arrow. They are thus different from weapons presumed to have ...

s, engraving tools, Sharp knife blades, and drilling and piercing tools. Humankind gradually evolved from early members of the genus ''

Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' (modern humans), plus several extinct species classified as either ancestral to or closely relat ...

''—such as ''Homo habilis'', who used simple stone tools—into

anatomically modern humans Early modern human (EMH) or anatomically modern human (AMH) are terms used to distinguish ''Homo sapiens'' (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans from extin ...

as well as behaviourally modern humans by the

Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coi ...

. During the end of the Paleolithic Age, specifically the Middle or Upper Paleolithic Age, humans began to produce the earliest works of art and to engage in

religious Religion is usually defined as a social- cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatural, ...

or spiritual behavior such as

burial Burial, also known as interment or inhumation, is a method of final disposition whereby a dead body is placed into the ground, sometimes with objects. This is usually accomplished by excavating a pit or trench, placing the deceased and objec ...

and

ritual A ritual is a sequence of activities involving gestures, words, actions, or objects, performed according to a set sequence. Rituals may be prescribed by the traditions of a community, including a religious community. Rituals are characterized, b ...

. Conditions during the Paleolithic Age went through a set of glacial and

interglacial period An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene ...

s in which the

climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologica ...

periodically fluctuated between warm and cool temperatures. Archaeological and genetic data suggest that the source populations of Paleolithic humans survived in sparsely-wooded areas and dispersed through areas of high primary productivity while avoiding dense forest-cover. By BP, the first humans set foot in Australia. By BP, humans lived at 61°N latitude in Europe. By BP, Japan was reached, and by BP humans were present in

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...

, above the Arctic Circle. By the end of the Upper Paleolithic Age humans had crossed

Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...

and expanded throughout the Americas.

Etymology

The term "

Palaeolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός '' palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...

" was coined by archaeologist John Lubbock in 1865. It derives from Greek: παλαιός, ''palaios'', "old"; and λίθος, ''lithos'', "stone", meaning "old age of the stone" or "Old Stone Age".

Paleogeography and climate

The Paleolithic coincides almost exactly with the

epoch of geologic time, which lasted from 2.6 million years ago to about 12,000 years ago. This epoch experienced important geographic and climatic changes that affected human societies. During the preceding

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58drift from possibly as far as from their present locations to positions only from their current location. South America became linked to North America through the

Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...

, bringing a nearly complete end to South America's distinctive

marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...

fauna. The formation of the isthmus had major consequences on global temperatures, because warm

equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can al ...

ial ocean currents were cut off, and the cold Arctic and Antarctic waters lowered temperatures in the now-isolated Atlantic Ocean. Most of

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

formed during the Pliocene to connect the continents of North and South America, allowing fauna from these continents to leave their native habitats and colonize new areas. Africa's collision with Asia created the Mediterranean, cutting off the remnants of the

Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

. During the

, the modern

continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...

s were essentially at their present positions; the

tectonic plate Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large t ...

s on which they sit have probably moved at most from each other since the beginning of the period. Climates during the Pliocene became cooler and drier, and seasonal, similar to modern climates.

Ice sheet In glaciology, an ice sheet, also known as a continental glacier, is a mass of glacial ice that covers surrounding terrain and is greater than . The only current ice sheets are in Antarctica and Greenland; during the Last Glacial Period at ...

s grew on

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...

. The formation of an Arctic ice cap around 3 million years ago is signaled by an abrupt shift in

oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as we ...

isotope Isotopes are two or more types of atoms that have the same atomic number (number of protons in their nuclei) and position in the periodic table (and hence belong to the same chemical element), and that differ in nucleon numbers ( mass number ...

ratios and ice-rafted cobbles in the North Atlantic and North

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...

beds. Mid-latitude

glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate bet ...

probably began before the end of the epoch. The global cooling that occurred during the Pliocene may have spurred on the disappearance of forests and the spread of

grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...

s and

savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...

s. The

climate was characterized by repeated glacial cycles during which

continental glacier In glaciology, an ice sheet, also known as a continental glacier, is a mass of glacial ice that covers surrounding terrain and is greater than . The only current ice sheets are in Antarctica and Greenland; during the Last Glacial Period at L ...

s pushed to the 40th parallel in some places. Four major glacial events have been identified, as well as many minor intervening events. A major event is a general glacial excursion, termed a "glacial". Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials". Each glacial advance tied up huge volumes of water in continental ice sheets deep, resulting in temporary sea level drops of or more over the entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions. Ice age fauna of northern Spain - Mauricio Antón

Ice age fauna of northern Spain - Mauricio Antón

The effects of glaciation were global.

was ice-bound throughout the Pleistocene and the preceding Pliocene. The

Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...

were covered in the south by the

Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and ...

n ice cap. There were glaciers in New Zealand and

Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...

. The now decaying glaciers of

Mount Kenya Mount Kenya ( Kikuyu: ''Kĩrĩnyaga'', Kamba, ''Ki Nyaa'') is the highest mountain in Kenya and the second-highest in Africa, after Kilimanjaro. The highest peaks of the mountain are Batian (), Nelion () and Point Lenana (). Mount Kenya is loc ...

Mount Kilimanjaro Mount Kilimanjaro () is a dormant volcano in Tanzania. It has three volcanic cones: Kibo, Mawenzi, and Shira. It is the highest mountain in Africa and the highest free-standing mountain above sea level in the world: above sea level and ab ...

, and the Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of

Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the Er ...

and to the west in the

Atlas mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through Moroc ...

. In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the Laurentide covered the east. The Fenno-Scandian ice sheet covered northern Europe, including Great Britain; the Alpine ice sheet covered the Alps. Scattered domes stretched across

and the Arctic shelf. The northern seas were frozen. During the late Upper Paleolithic (Latest Pleistocene) BP, the

land bridge between Asia and North America was blocked by ice, which may have prevented early Paleo-Indians such as the

Clovis culture The Clovis culture is a prehistoric Paleoamerican culture, named for distinct stone and bone tools found in close association with Pleistocene fauna, particularly two mammoths, at Blackwater Locality No. 1 near Clovis, New Mexico, in 1936 ...

from directly crossing

to reach the Americas. According to Mark Lynas (through collected data), the Pleistocene's overall climate could be characterized as a continuous

El Niño El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date ...

with

trade winds The trade winds or easterlies are the permanent east-to-west prevailing winds that flow in the Earth's equatorial region. The trade winds blow mainly from the northeast in the Northern Hemisphere and from the southeast in the Southern Hemisp ...

in the south

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

weakening or heading east, warm air rising near

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

, warm water spreading from the west Pacific and the

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by ...

to the east Pacific, and other El Niño markers. The Paleolithic is often held to finish at the end of the ice age (the end of the Pleistocene epoch), and Earth's climate became warmer. This may have caused or contributed to the extinction of the

Pleistocene megafauna Pleistocene megafauna is the set of large animals that lived on Earth during the Pleistocene epoch. Pleistocene megafauna became extinct during the Quaternary extinction event resulting in substantial changes to ecosystems globally. The role of hu ...

, although it is also possible that the late

Pleistocene extinctions The Quaternary period (from 2.588 ± 0.005 million years ago to the present) has seen the extinctions of numerous predominantly megafaunal species, which have resulted in a collapse in faunal density and diversity and the extinction of key ecolog ...

were (at least in part) caused by other factors such as disease and overhunting by humans. New research suggests that the extinction of the

woolly mammoth The woolly mammoth (''Mammuthus primigenius'') is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus sub ...

may have been caused by the combined effect of climatic change and human hunting. Scientists suggest that climate change during the end of the Pleistocene caused the mammoths' habitat to shrink in size, resulting in a drop in population. The small populations were then hunted out by Paleolithic humans. The global warming that occurred during the end of the Pleistocene and the beginning of the

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...

may have made it easier for humans to reach mammoth habitats that were previously frozen and inaccessible. Small populations of woolly mammoths survived on isolated Arctic islands, Saint Paul Island and

Wrangel Island Wrangel Island ( rus, О́стров Вра́нгеля, r=Ostrov Vrangelya, p=ˈostrəf ˈvrangʲɪlʲə; ckt, Умӄиԓир, translit=Umqiḷir) is an island of the Chukotka Autonomous Okrug, Russia. It is the 91st largest island in the w ...

, until BP and BP respectively. The Wrangel Island population became extinct around the same time the island was settled by prehistoric humans. There is no evidence of prehistoric human presence on Saint Paul island (though early human settlements dating as far back as 6500 BP were found on the nearby

Aleutian Islands The Aleutian Islands ( ; ; ale, Unangam Tanangin, "land of the Aleuts"; possibly from the Chukchi ''aliat'', or "island")—also called the Aleut Islands, Aleutic Islands, or, before 1867, the Catherine Archipelago—are a chain of 14 main, ...

Human way of life

Nearly all of our knowledge of Paleolithic human culture and way of life comes from

archaeology Archaeology or archeology is the scientific study of human activity through the recovery and analysis of material culture. The archaeological record consists of Artifact (archaeology), artifacts, architecture, biofact (archaeology), biofacts ...

and

ethnographic Ethnography (from Greek ''ethnos'' "folk, people, nation" and ''grapho'' "I write") is a branch of anthropology and the systematic study of individual cultures. Ethnography explores cultural phenomena from the point of view of the subject o ...

comparisons to modern hunter-gatherer cultures such as the !Kung San who live similarly to their Paleolithic predecessors. The economy of a typical Paleolithic society was a hunter-gatherer economy. Humans hunted wild animals for meat and gathered food, firewood, and materials for their tools, clothes, or shelters. Human population density was very low, around only . This was most likely due to low body fat,

infanticide Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of resou ...

, high levels of physical activity among women, late weaning of infants, and a

nomad A nomad is a member of a community without fixed habitation who regularly moves to and from the same areas. Such groups include hunter-gatherers, pastoral nomads (owning livestock), tinkers and trader nomads. In the twentieth century, the po ...

ic lifestyle. Like contemporary hunter-gatherers, Paleolithic humans enjoyed an abundance of leisure time unparalleled in both

Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several part ...

farming societies and modern industrial societies. At the end of the Paleolithic, specifically the Middle or Upper Paleolithic, humans began to produce works of art such as

cave painting In archaeology, Cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric origin, and the oldest known are more than 40,00 ...

s, rock art and

jewellery Jewellery ( UK) or jewelry ( U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a we ...

and began to engage in religious behavior such as burials and rituals.

Distribution

At the beginning of the Paleolithic, hominins were found primarily in eastern Africa, east of the

Great Rift Valley The Great Rift Valley is a series of contiguous geographic trenches, approximately in total length, that runs from Lebanon in Asia to Mozambique in Southeast Africa. While the name continues in some usages, it is rarely used in geology as ...

. Most known hominin fossils dating earlier than one million years before present are found in this area, particularly in

Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...

, and

. By BP, groups of hominins began leaving Africa and settling southern Europe and Asia. Southern Caucasus was occupied by BP, and northern China was reached by BP. By the end of the Lower Paleolithic, members of the hominin family were living in what is now China, western Indonesia, and, in Europe, around the Mediterranean and as far north as England, France, southern Germany, and Bulgaria. Their further northward expansion may have been limited by the lack of control of fire: studies of cave settlements in Europe indicate no regular use of fire prior to BP. East Asian fossils from this period are typically placed in the genus ''

Homo erectus ''Homo erectus'' (; meaning " upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as ''H. heidelbergensis'' and ''H. antecessor' ...

''. Very little fossil evidence is available at known Lower Paleolithic sites in Europe, but it is believed that hominins who inhabited these sites were likewise ''Homo erectus''. There is no evidence of hominins in America, Australia, or almost anywhere in Oceania during this time period. Fates of these early colonists, and their relationships to modern humans, are still subject to debate. According to current archaeological and genetic models, there were at least two notable expansion events subsequent to peopling of Eurasia BP. Around 500,000 BP a group of early humans, frequently called ''

Homo heidelbergensis ''Homo heidelbergensis'' (also ''H. sapiens heidelbergensis''), sometimes called Heidelbergs, is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of '' H. erectus'' ...

'', came to Europe from Africa and eventually evolved into ''Homo neanderthalensis'' (

Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While ...

s). In the Middle Paleolithic, Neanderthals were present in the region now occupied by Poland. Both ''Homo erectus'' and ''Homo neanderthalensis'' became extinct by the end of the Paleolithic. Descended from ''Homo sapiens'', the anatomically modern ''

Homo sapiens sapiens Human taxonomy is the classification of the human species (systematic name ''Homo sapiens'', Latin: "wise man") within zoological taxonomy. The systematic genus, ''Homo'', is designed to include both anatomically modern humans and extinct va ...

'' emerged in eastern Africa BP, left Africa around 50,000 BP, and expanded throughout the planet. Multiple hominid groups coexisted for some time in certain locations. ''Homo neanderthalensis'' were still found in parts of Eurasia BP years, and engaged in an unknown degree of interbreeding with ''Homo sapiens sapiens''. DNA studies also suggest an unknown degree of interbreeding between ''Homo sapiens sapiens'' and '' Homo sapiens denisova''. Hominin fossils not belonging either to ''Homo neanderthalensis'' or to ''Homo sapiens'' species, found in the Altai Mountains and Indonesia, were radiocarbon dated to BP and BP respectively. For the duration of the Paleolithic, human populations remained low, especially outside the equatorial region. The entire population of Europe between 16,000 and 11,000 BP likely averaged some 30,000 individuals, and between 40,000 and 16,000 BP, it was even lower at 4,000–6,000 individuals. However, remains of thousands of butchered animals and tools made by Palaeolithic humans were found in Lapa do Picareiro ( pt), a cave in

Portugal Portugal, officially the Portuguese Republic, In recognized minority languages of Portugal: :* mwl, República Pertuesa is a country located on the Iberian Peninsula, in Southwestern Europe, and whose territory also includes the Macaronesian ...

, dating back between 41,000 and 38,000 years ago.

Technology

Tools

Paleolithic humans made tools of stone, bone (primarily deer), and wood. The early paleolithic hominins, ''

Australopithecus ''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Austral ...

'', were the first users of stone tools. Excavations in

Gona, Ethiopia Gona is an archaeological site in the Afar Triangle of Ethiopia located in the Ethiopian Lowlands. The site, near the Middle Awash and Hadar regions, is primarily known for paleoanthropological study, including excavations of Late Miocene a ...

have produced thousands of artifacts, and through radioisotopic dating and

magnetostratigraphy Magnetostratigraphy is a geophysical correlation technique used to date sedimentary and volcanic sequences. The method works by collecting oriented samples at measured intervals throughout the section. The samples are analyzed to determine their ' ...

, the sites can be firmly dated to 2.6 million years ago. Evidence shows these early hominins intentionally selected raw stone with good flaking qualities and chose appropriate sized stones for their needs to produce sharp-edged tools for cutting. The earliest Paleolithic stone tool industry, the

Oldowan The Oldowan (or Mode I) was a widespread stone tool archaeological industry (style) in prehistory. These early tools were simple, usually made with one or a few flakes chipped off with another stone. Oldowan tools were used during the Lower ...

, began around 2.6 million years ago. It produced tools such as choppers, burins, and

stitching awl A stitching awl is a tool with which holes can be punctured in a variety of materials, or existing holes can be enlarged. It is also used for sewing heavy materials, such as leather or canvas. It is a thin, tapered metal shaft, coming to a shar ...

s. It was completely replaced around 250,000 years ago by the more complex

Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped " hand axes" associat ...

industry, which was first conceived by ''

Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unre ...

'' around 1.8–1.65 million years ago. The Acheulean implements completely vanish from the archaeological record around 100,000 years ago and were replaced by more complex Middle Paleolithic tool kits such as the

Mousterian The Mousterian (or Mode III) is an archaeological industry of stone tools, associated primarily with the Neanderthals in Europe, and to the earliest anatomically modern humans in North Africa and West Asia. The Mousterian largely defines the ...

and the

Aterian The Aterian is a Middle Stone Age (or Middle Palaeolithic) stone tool industry centered in North Africa, from Mauritania to Egypt, but also possibly found in Oman and the Thar Desert. The earliest Aterian dates to c. 150,000 years ago, at th ...

industries. Lower Paleolithic humans used a variety of stone tools, including

hand axe A hand axe (or handaxe or Acheulean hand axe) is a prehistoric stone tool with two faces that is the longest-used tool in human history, yet there is no academic consensus on what they were used for. It is made from stone, usually flint or che ...

s and choppers. Although they appear to have used hand axes often, there is disagreement about their use. Interpretations range from cutting and chopping tools, to digging implements, to flaking cores, to the use in traps, and as a purely ritual significance, perhaps in courting behavior. William H. Calvin has suggested that some hand axes could have served as "killer Frisbees" meant to be thrown at a herd of animals at a waterhole so as to stun one of them. There are no indications of

hafting Hafting is a process by which an artifact, often bone, stone, or metal is attached to a ''haft'' (handle or strap). This makes the artifact more useful by allowing it to be shot (arrow), thrown by hand (spear), or used with more effective levera ...

, and some artifacts are far too large for that. Thus, a thrown hand axe would not usually have penetrated deeply enough to cause very serious injuries. Nevertheless, it could have been an effective weapon for defense against predators. Choppers and scrapers were likely used for skinning and butchering scavenged animals and sharp-ended sticks were often obtained for digging up edible roots. Presumably, early humans used wooden spears as early as 5 million years ago to hunt small animals, much as their relatives, chimpanzees, have been observed to do in

Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣� ...

, Africa. Lower Paleolithic humans constructed shelters, such as the possible wood hut at Terra Amata.

Fire use

Fire was used by the Lower Paleolithic hominins ''

'' and ''

'' as early as 300,000 to 1.5 million years ago and possibly even earlier by the early Lower Paleolithic (Oldowan) hominin ''

Homo habilis ''Homo habilis'' ("handy man") is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.31 million years ago to 1.65 million years ago (mya). Upon species description in 1964, ''H. habilis'' was highly ...

'' or by robust ''

Australopithecine Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' ( cladistically including the genera ''Homo'', ''Paranthropus'', and '' Kenyanthropus''), and it typically include ...

s'' such as ''

Paranthropus ''Paranthropus'' is a genus of extinct hominin which contains two widely accepted species: '' P. robustus'' and '' P. boisei''. However, the validity of ''Paranthropus'' is contested, and it is sometimes considered to be synonymous with '' Au ...

''. However, the use of fire only became common in the societies of the following

Middle Stone Age The Middle Stone Age (or MSA) was a period of African prehistory between the Early Stone Age and the Late Stone Age. It is generally considered to have begun around 280,000 years ago and ended around 50–25,000 years ago. The beginnings of ...

and

Middle Paleolithic The Middle Paleolithic (or Middle Palaeolithic) is the second subdivision of the Paleolithic or Old Stone Age as it is understood in Europe, Africa and Asia. The term Middle Stone Age is used as an equivalent or a synonym for the Middle Paleo ...

. Use of fire reduced mortality rates and provided protection against predators. Early hominins may have begun to cook their food as early as the Lower Paleolithic ( million years ago) or at the latest in the early Middle Paleolithic ( years ago). Some scientists have hypothesized that hominins began cooking food to defrost frozen meat, which would help ensure their survival in cold regions. Archaeologists cite morphological shifts in cranial anatomy as evidence for emergence of cooking and

food processing Food processing is the transformation of agricultural products into food, or of one form of food into other forms. Food processing includes many forms of processing foods, from grinding grain to make raw flour to home cooking to complex industr ...

technologies. These morphological changes include decreases in molar and jaw size, thinner tooth enamel, and decrease in gut volume During much of the

epoch, our ancestors relied on simple

techniques such as

roasting Roasting is a cooking method that uses dry heat where hot air covers the food, cooking it evenly on all sides with temperatures of at least from an open flame, oven, or other heat source. Roasting can enhance the flavor through caramelizatio ...

The

Upper Palaeolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coi ...

saw the emergence of boiling, an advance in

technology which rendered plant foods more digestible, decreased their toxicity, and maximised their nutritional value Thermally altered rock (heated stones) are easily identifiable in the archaeological record. Stone-boiling and pit-baking were common techniques which involved heating large pebbles then transferring the hot stones into a perishable container to heat the water This technology is typified in the

Middle Palaeolithic The Middle Paleolithic (or Middle Palaeolithic) is the second subdivision of the Paleolithic or Old Stone Age as it is understood in Europe, Africa and Asia. The term Middle Stone Age is used as an equivalent or a synonym for the Middle Paleo ...

example of the

Abri Pataud L'Abri Pataud, or the Pataud Shelter in English, is a prehistoric site found in the middle of the village Les Eyzies-de-Tayac-Sireuil in Dordogne, Aquitaine, southwestern France. The site includes human remains, stone tools, and early cultural art ...

hearths

Raft

The Lower Paleolithic ''Homo erectus'' possibly invented

raft A raft is any flat structure for support or transportation over water. It is usually of basic design, characterized by the absence of a hull. Rafts are usually kept afloat by using any combination of buoyant materials such as wood, sealed barrel ...

s ( BP) to travel over large bodies of water, which may have allowed a group of ''Homo erectus'' to reach the island of Flores and evolve into the small hominin ''