Pale-tailed Barbthroat
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The pale-tailed barbthroat (''Threnetes leucurus'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
in the family Trochilidae. It is found throughout much of the
Amazon Basin The Amazon basin is the part of South America drained by the Amazon River and its tributary, tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries ...
from the eastern Andean foothills to the Atlantic Ocean.


Taxonomy and systematics

The pale-tailed barbthroat was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1766 in the
twelfth edition Twelfth can mean: *The Twelfth Amendment to the United States Constitution *The Twelfth, a Protestant celebration originating in Ireland In mathematics: * 12th, an ordinal number; as in the item in an order twelve places from the beginning, follo ...
of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Trochilus leucurus''. Linnaeus based his short description on the "white-tailed humming-bird" that had been described and illustrated by the English naturalist George Edwards in 1758. The type locality is
Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...
. The specific epithet is from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''leukouros'' meaning "white-tailed". The pale-tailed barbthroat is now placed in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Threnetes'' that was introduced in 1852 by the English ornithologist
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist who published monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould (illustrator), Elizabeth Gould, and several other artists, includ ...
. Four
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised: * ''T. l. cervinicauda''
Gould Gould may refer to: People * Gould (name), a surname Places United States * Gould, Arkansas, a city * Gould, Colorado, an unincorporated community * Gould, Ohio, an unincorporated community * Gould, Oklahoma, a town * Gould, West Virginia, an ...
, 1855 * ''T. l. rufigastra''
Cory As a given name, Cory is used by both males and females. It is a variation of the name Cora, meaning "(the) Maiden", which is a title of the goddess Persephone. The name also can have origins from the Gaelic word ''coire'', which means "in a cauldr ...
, 1915 * ''T. l. leucurus'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1766) * ''T. l. medianus''
Hellmayr Carl Eduard Hellmayr (29 January 1878 – 24 February 1944) was an Austrian ornithologist. Biography Hellmayr was born in Vienna and studied at the University of Vienna, although he did not complete his degree. After his studies he worked in Vie ...
, 1929 The taxonomy of the pale-tailed and sooty barbthroat (''T. niger'') complex is unsettled. The two were originally described as separate species, then combined under the name pale-tailed barbthroat but with the
binomial Binomial may refer to: In mathematics *Binomial (polynomial), a polynomial with two terms *Binomial coefficient, numbers appearing in the expansions of powers of binomials *Binomial QMF, a perfect-reconstruction orthogonal wavelet decomposition * ...
''T. niger'', which had priority. In the early 2000s they were again split, with the two species recognized by the
International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...
(IOC), the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
, and
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' (HBW). However, the assignment of subspecies between them varies. The IOC and Clements assign the above four subspecies to the pale-tailed barbthroat and two others to the sooty barbthroat. HBW transfers subspecies ''loehkeni'' from the sooty to the pale-tailed, leaving the former
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
. Other authors suggest that ''loehkeni'' should be treated as a separate species.Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021del Hoyo, J., N. Collar, and G. M. Kirwan (2020). Pale-tailed Barbthroat (''Threnetes leucurus''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.patbar1.01 retrieved November 15, 2021 The pale-tailed barbthroat and the other two members of its genus, band-tailed barbthroat (''T. ruckeri'') and sooty barbthroat (''T. niger''), form a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved August 24, 2021


Description

The pale-tailed barbthroat is long. Males weigh and females . The adult male has bronze-green upperparts and breast, a dark ear patch and throat with a reddish patch below the later, and a pale malar stripe. The tail is dark with various shades and extent of color on the outer feathers. The belly is gray with ochre tints. It has a nearly straight bill. As with other
hermit A hermit, also known as an eremite (adjectival form: hermitic or eremitic) or solitary, is a person who lives in seclusion. Eremitism plays a role in a variety of religions. Description In Christianity, the term was originally applied to a Chr ...
hummingbirds, the sexes are similar; the female's bill is somewhat more decurved than the male's and the plumage has less contrast among the throat, breast, and belly. Young birds resemble the adult but have ochraceous feather edges. The subspecies differ somewhat in the tail and belly coloration, and there are intergrades between ''cervinicauda'' and ''rufigastra'' and between ''rufigastra'' and ''leucurus''.


Distribution and habitat

The subspecies of pale-tailed barbthroat are distributed thus: * ''T. l. cervinicauda'', eastern Colombia south through eastern Ecuador into northeastern Peru and east into western Amazonian Brazil * ''T. l. rufigastra'', eastern Peru south of the
Marañón River The Marañón River (, , ) is the principal or mainstem source of the Amazon River, arising about 160 km (100 miles) to the northeast of Lima, Peru, and flowing northwest across plateaus 3,650 m (12,000 feet) high, it runs through a deeply ero ...
to northern Bolivia * ''T. l. leucurus'', southern and eastern Venezuela through Guyana, Suriname, and Amazonian Brazil to northeastern Bolivia (but not confirmed in French Guiana)Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved August 24, 2021 * ''T. l. medianus'', northeastern Amazonian Brazil south of the Amazon The pale-tailed barbthroat inhabits open and semi-open spaces within humid lowland and higher
tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...
, and other landscapes such as
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, '' várzea'' and ''
igapó (, from Tupi language, Old Tupi: "root forest") is a word used in Brazil for Blackwater river, blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur ...
'' swamp forests, and plantations. It is found from sea level to in Venezuela, in Peru mainly to but as high as , and in Ecuador mainly to and as high as .


Behavior


Movement

The pale-tailed barbthroat is presumed to be a year round resident throughout its range.


Feeding

Like other hermit hummingbirds, the pale-tailed barbthroat is a " trap-line" feeder, visiting a circuit of flowering plants. It feeds on nectar at ''
Heliconia ''Heliconia'' is a genus of flowering plants in the monotypic family Heliconiaceae. Most of the 194 known species are native to the tropical Americas, but a few are indigenous to certain islands of the western Pacific and Maluku (province), ...
'' and several other tubular flowers and also on small
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s.


Breeding

The pale-tailed barbthroat's nesting season varies widely across its range. Its nest is a cone-shaped cup of plant and other fibers and cobweb, covered with lichens, and attached to the underside tip of a long drooping leaf such as a palm frond. The female alone incubates the white eggs.


Vocalization

The pale-tailed barbthroat's song is "a fast high-pitched phrase of 5–10 notes repeated at intervals of several seconds". Its call is "a high-pitched sharp 'tseet', sometimes doubled."


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the pale-tailed barbthroat as being of Least Concern. It has a very large range, and though its population has not been quantified it is believed to be stable. It is considered local and uncommon throughout its range and occurs in several protected areas.


References


External links

* (for Suriname)
Pale-tailed Barbthroat photo gallery
VIREO {{Taxonbar, from=Q1270356 pale-tailed barbthroat Hummingbird species of South America Birds of the Guiana Shield Birds of the Amazon rainforest pale-tailed barbthroat pale-tailed barbthroat Birds of Brazil Taxonomy articles created by Polbot