Pale-legged Hornero
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The pale-legged hornero (''Furnarius leucopus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is found in
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...
, and
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
.


Taxonomy and systematics

The pale-legged hornero's taxonomy is unsettled. The
International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...
(IOC),
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' (HBW), and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
assign it these four subspecies:HBW and BirdLife International (2022) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 7. Available at: https://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v7_Dec22.zip retrieved December 13, 2022Clements, J. F., P.C. Rasmussen, T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, A. Spencer, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2023. The eBird/Clements checklist of birds of the world: v2023. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved October 28, 2023 *''F. l. leucopus'' Swainson, 1838 *''F. l. tricolor'' Giebel, 1868 *''F. l. araguaiae''
Pinto Pinto is a Portuguese, Spanish, Jewish (Sephardic), and Italian surname. It is a high-frequency surname in all Portuguese-speaking countries and is also widely present in Spanish-speaking countries, Italy, India (especially in Mangalore, Karnata ...
& Camargo, 1952
*''F. l. assimilis''
Cabanis Cabanis is a surname. Notable people with the surname include: * Bernard Cabanis (born 1950), French ice hockey player * George Cabanis (1815–1892), American politician * James Cabanis (1838–1920), American politician, son of George Cabanis *Jea ...
&
Heine Heine is both a surname and a given name of German origin. People with that name include: People with the surname * Albert Heine (1867–1949), German actor * Alice Heine (1858–1925), American-born princess of Monaco * Armand Heine (1818–1883) ...
, 1860
The South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
(SACC) adds three others, ''F. l. cinnamomeus'' (
Lesson A lesson or class is a structured period of time where learning is intended to occur. It involves one or more students (also called pupils or learners in some circumstances) being taught by a teacher or instructor. A lesson may be either one ...
, 1844), ''F. l. longirostris'' ( von Pelzeln, 1856), and ''F. l. endoecus'' (
Cory As a given name, Cory is used by both males and females. It is a variation of the name Cora, meaning "(the) Maiden", which is a title of the goddess Persephone. The name also can have origins from the Gaelic word ''coire'', which means "in a cauldr ...
, 1919).Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 28 September 2023. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved October 20, 2023 The IOC, HBW, and Clements treat ''cinnamomeus'' as the species Pacific hornero and the other two as the
Caribbean hornero The Caribbean hornero (''Furnarius longirostris'') is a species of bird in the Furnariinae subfamily of the ovenbird family (biology), family Furnariidae. It is found in Colombia and Venezuela. Taxonomy and systematics The Caribbean hornero' ...
. Early authors (e.g. Chapman) had treated them separately.Chapman, F. M. (1926). The distribution of bird-life in Ecuador: a contribution to a study of the origin of Andean bird-life. Bulletin of the American Museum of Natural History 55:1–784. The SACC accepts that ''cinnamomeus'' may deserve species rank but declined to make the split due to "insufficient published data". Some authors have treated what is now the pale-billed hornero (''F. torridus'') as a subspecies of the pale-legged hornero.Kirwan, G. M., J. del Hoyo, J. V. Remsen, Jr., N. Collar, and E. de Juana (2021). Pale-legged Hornero (''Furnarius leucopus''), version 2.0. In Birds of the World (B. K. Keeney, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.palhor2.02 retrieved August 22, 2023 This article follows the four-subspecies model.


Description

The pale-legged hornero is long and weighs about . It is a medium-sized hornero with a long and nearly straight bill. The sexes' plumages are alike. Adults of the nominate subspecies ''F. l. leucopus'' have a wide whitish
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
, brownish gray ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
, and a tawny-rufous malar area. Their crown is dark
rufescent Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
brown. Their back, rump, and uppertail coverts average bright orange rufous but vary between tawny rufous and rufous amber. Their tail is chestnut. Their wing coverts are chestnut and their flight feathers blackish with a wide chestnut band. Their throat is white that becomes tawny-
ochraceous Ochre ( ; , ), iron ochre, or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the colou ...
on the breast. Their flanks are paler tawny-ochraceous, the center to their belly nearly whitish, and their undertail coverts whitish with dark brown bases. Their iris is usually reddish brown or chestnut, and gray-brown or gray in ''tricolor''. Their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
is dusky horn at its base with a paler culmen and tip and their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
is also pale. Their legs and feet are pale pinkish, pearly gray, or whitish. Juveniles resemble adults but for a visibly shorter bill and the shape of their flight and tail feathers. Subspecies ''F. l. tricolor'' has a grayer crown, more ochraceous back, and paler wings and tail than the nominate. ''F. l. assimilis'' has a lighter, more ochraceous rump, wings, and tail than ''tricolor'', and a paler wing band. ''F. l. araguaiae'' is intermediate between ''tricolor'' and ''assimilis'', with a brighter back than the former and a smaller wing band.


Distribution and habitat

The subspecies of the pale-legged hornero are found thus: *''F. l. leucopus'': along the rios
Negro In the English language, the term ''negro'' (or sometimes ''negress'' for a female) is a term historically used to refer to people of Black people, Black African heritage. The term ''negro'' means the color black in Spanish and Portuguese (from ...
and Branco in northern Brazil and in southwestern Guyana *''F. l. tricolor'': eastern Peru, western Brazil east into
Pará Pará () is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian st ...
, and into Bolivia to Santa Cruz Department; one record in southeastern Ecuador *''F. l. araguaiae'': western
Tocantins Tocantins () is one of the 26 states of Brazil. It is the newest state, formed in 1988 and encompassing what had formerly been the northern two-fifths of the state of Goiás. Tocantins covers and had an estimated population of 1,496,880 in 2014 ...
and eastern
Mato Grosso Mato Grosso ( – ) is one of the states of Brazil, the List of Brazilian states by area, third largest by area, located in the Central-West Region, Brazil, Central-West region. The state has 1.66% of the Brazilian population and is responsible ...
states in central Brazil along the rios Araguaia and das Mortes *''F. l. assimilis'': eastern and southern Brazil between
Maranhão Maranhão () is a States of Brazil, state in Brazil. Located in the country's Northeast Region, Brazil, Northeast Region, it has a population of about 7 million and an area of and it is divided into 217 municipalities. Clockwise from north, it ...
,
Pernambuco Pernambuco ( , , ) is a States of Brazil, state of Brazil located in the Northeast Region, Brazil, Northeast region of the country. With an estimated population of 9.5 million people as of 2024, it is the List of Brazilian states by population, ...
, and
Mato Grosso do Sul Mato Grosso do Sul ( ) is one of Federative units of Brazil, Brazil's 27 federal units, located in the southern part of the Central-West Region, Brazil, Central-West Region, bordering five Brazilian states: Mato Grosso (to the north), Goiás and ...
, and extreme southeastern Bolivia The pale-legged hornero inhabits a wide variety of semi-open to open landscapes. These include forest and woodlands along rivers (
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
), the edges of
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
, agricultural areas, and parks and gardens in towns. It favors humid areas, usually near water. In elevation it mostly occurs below , though it seldom exceeds in Bolivia and locally reaches in Peru.


Behavior


Movement

The pale-legged hornero is essentially a year-round resident throughout its range. The record in Ecuador and a few others outside its usual range could be "more or less local movements, sometimes following rivers, or otherwise overlooked populations?".


Feeding

The pale-legged hornero's diet is mostly a variety of
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s. It also includes other small invertebrates like snails and there is one record of an individual eating a toad. It forages singly or in pairs while walking on the ground, turning over leaves to glean its prey.


Breeding

The pale-legged hornero's breeding biology is " rprisingly poorly known for such a relatively abundant, noisy, and conspicuous species". Its breeding season has not been defined. Though its nest has not been formally described, it is an "oven" of clay and animal dung with an inner chamber lined with dry plant matter. Both members of a pair construct it, typically on a tree branch but also on horizontal structures like the crossbars of utility poles. The clutch size is not known. The incubation period at one nest in Peru was 16 to 17 days and the time to fledging 26 to 33 days. Both parents provisioned the nestlings.


Vocalization

The pale-legged hornero's song is a "long series of loud, explosive, piercing, staccato notes, which decelerates and descends in pitch...rendered ''PIPIPI’PI’pi’pi-pi-pi-pi-pi pee pee pu''". Its calls are "a loud, rich, scratchy or reedy ''chet'' or ''kyeek'', or a descending ''cheeop''".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the pale-legged hornero as being of Least Concern. It has a very large range, and though its population size is not known it is believed to be increasing. No immediate threats have been identified. It is considered uncommon to very common. It " nefits from moderate anthropogenic habitat alteration, and...it has extended its range into deforested areas". It occurs in several protected areas.


References

{{Taxonbar, from=Q1265893 pale-legged hornero Birds of the Amazon rainforest Birds of the Bolivian Amazon Birds of Peru Birds of Brazil Birds of the Caatinga pale-legged hornero Taxonomy articles created by Polbot Taxa named by William Swainson