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The pale-browed treehunter (''Cichlocolaptes leucophrus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the Furnariinae subfamily of the ovenbird
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Furnariidae. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.


Taxonomy and systematics

The pale-browed treehunter's taxonomy is unsettled. The South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
, the
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
, and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
treat it as a species with two subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''C. l. leucophrus'' (
Jardine Jardine is a surname. Notable people with the surname include: * Al Jardine (born 1942), member of the Beach Boys * Alexander Jardine (Medal of Honor) (1874–1949), American Medal of Honor recipient * Antonio Jardine (born 1988), NCAA college ba ...
&
Selby Selby is a market town and civil parish in the Selby District of North Yorkshire, England, south of York on the River Ouse, with a population at the 2011 census of 14,731. The town was historically part of the West Riding of Yorkshire until ...
, 1830) and ''C. l. holti'' (
Pinto Pinto is a Portuguese, Spanish, Jewish (Sephardic), and Italian surname. It is a high-frequency surname in all Portuguese-speaking countries and is also widely present in Spanish-speaking countries, Italy, India especially in Mangalore, Kar ...
, 1941).
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' (HBW) treats the two taxa as separate species, the large and small pale-browed treehunters respectively.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 May 2023. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved May 31, 2023Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022 The pale-browed treehunter shares genus ''Cichlocolaptes'' with the
cryptic treehunter The cryptic treehunter (''Cichlocolaptes mazarbarnetti'') is or was a species of bird in the Furnariinae subfamily of the ovenbird family Furnariidae. Its status, and even its existence as a species, are disputed. It is or was endemic to Brazi ...
(''C. mazarbarnetti''), which is believed to be extinct. This article follows the one-species two-subspecies model.


Description

The nominate subspecies of the pale-browed treehunter is about long; subspecies ''C. l. holti'' is about long. Both subspecies have a long dagger-like bill, and male and female plumages are alike. Adults of the nominate subspecies have a buffish white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
, blackish and dull buff
lores Lores may refer to: * Lore (anatomy) * Lores (surname) Lores is a surname. Notable people with the surname include: *Enrique Lores (born 1964/65), Spanish business executive *Horacio Lores, Argentine politician *Julio Lores (1908–1947), Peruvi ...
, and a dark brown band behind the eye on an otherwise duff buff face. Their crown is dark brown whose feathers have obvious buff to rusty-buff shafts. They have an indistinct blackish brown collar with buff streaks. Their back is
rufescent Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
brown with long blurry buff streaks and their rump and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
are a slightly paler brown with indistinct buff streaks. Their tail is pale rufous or cinnamon. Their wing coverts are brown with indistinct buff streaks and their flight feathers rich brown. Their throat is pale yellowish buff, their breast has strong pale buff and brown streaks, and their belly, flanks, and undertail coverts are like the breast with the streaks progressively fading. Their iris is very dark, their
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
dusky gray, their
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bon ...
a paler brown with gray towards the tip, and their legs and feet variable from greenish gray to pale brownish. Juveniles are much like adults but with less distinct streaking on their underparts. Compared to the nominate subspecies, ''C. l. holti'' is smaller, has a wider and paler supercilium, blacker feathers in the crown, slightly more olive upperparts, a darker rufous tail, and a plain buff throat.del Hoyo, J., J. V. Remsen, Jr., G. M. Kirwan, and N. Collar (2020). Pale-browed Treehunter (''Cichlocolaptes leucophrus''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.pabtre1.01 retrieved August 26, 2023


Distribution and habitat

The pale-browed treehunter is found in southeastern Brazil. The nominate subspecies occurs from southern
Bahia Bahia ( , , ; meaning "bay") is one of the 26 states of Brazil, located in the Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo, Minas Gerais, and Rio de Janeiro) and the 5th-largest ...
south to
Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the Rio de Janeiro (state), state of the same name, Brazil's List of Brazilian states by population, third-most populous state, and the List of largest citi ...
state. ''C. l. holti'' occurs from
São Paulo São Paulo (, ; Portuguese for 'Saint Paul') is the most populous city in Brazil, and is the capital of the state of São Paulo, the most populous and wealthiest Brazilian state, located in the country's Southeast Region. Listed by the Ga ...
state south to northeastern Rio Grande do Sul. The species inhabits humid lowland and montane
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperat ...
. The nominate subspecies occurs from sea level to while ''C. l. holti'' reaches only about .


Behavior


Movement

The pale-browed treehunter is a year-round resident throughout its range.


Feeding

The pale-browed treehunter feeds on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s. It often, and perhaps usually, forages as a member of a
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These are ...
. It feeds at any level of the forest between its understory and the subcanopy. It forages along larger limbs and branches, probing and gleaning for prey, with a strong preference for seeking it in
bromeliad The Bromeliaceae (the bromeliads) are a family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the tropical Americas, with several species found in the American subtropics and one in tropical west Africa ...
s and other epiphytes. It sometimes goes completely into clumps of bromeliads and may hang upside-down to feed.


Breeding

Nothing is known about the pale-browed treehunter's breeding biology.


Vocalization

The songs of the two subspecies of pale-browed treehunter differ slightly in pitch. In general they are "loud, ringing 'reep' notes, variable in number, sometimes interspersed with fast chattering notes". Their call is "a loud 'krip, shrip' ".


Status

The IUCN follows HBW taxonomy and so has separately assessed the "large" and "small" pale-browed treehunters. The assessements are the same: Both taxa are of Least Concern. They have large ranges and unknown population sizes that are believed to be decreasing. No immediate threats to either have been identified. Both subspecies are considered rare to locally common and each is found in several protected areas. The
Atlantic Forest The Atlantic Forest ( pt, Mata Atlântica) is a South American forest that extends along the Atlantic coast of Brazil from Rio Grande do Norte state in the northeast to Rio Grande do Sul state in the south and inland as far as Paraguay and t ...
has undergone extensive clearing, leaving little suitable habitat for them.


References

{{Taxonbar, from=Q616230 Cichlocolaptes Birds of the Atlantic Forest Endemic birds of Brazil Birds described in 1830 Taxonomy articles created by Polbot Taxa named by Sir William Jardine Taxa named by Prideaux John Selby