''Cornus nuttallii'', the Pacific dogwood, western dogwood, or mountain dogwood, is a species of

dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods or cornels, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous ...

native to western

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. The tree's name used by Hul'q'umi'num'-speaking nations is ''Kwi’txulhp''.

Description

It is a small to medium-sized

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

, reaching tall, often with a canopy spread of . Its

habit A habit (or wont, as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously. A 1903 paper in the '' American Journal of Psychology'' defined a "habit, from the standpoint of psychology, ...

varies based on the level of sunlight; in full sun it will have a short trunk with a crown as wide as it is tall, while under a canopy it will have a tapered trunk with a short, slender crown. The trunk attains in diameter. The

bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...

is reddish brown. The branches have fine hairs and the young bark is thin and smooth, becoming scale-like with ridges as it ages. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are opposite, simple, oval, long, and broad. They are green with stiff, appressed hairs on top, and hairier and lighter on the bottom. They turn orange to purplish in autumn. The

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s are individually small and inconspicuous, across, produced in a dense, rounded, greenish-white flower head in diameter; the 4–8 large white 'petals' are actually

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

s, each bract long and broad, creating the appearance of a larger flower head. The flowers commonly bloom twice per season, once in the spring and again in late summer or early fall. Appearing in September or October, the

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is a compound pink-red or orange

drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...

about long. They are produced in clusters containing 20–40 drupelets, each of which contains two seeds. The fruits are unpalatable.

Similar species

The eastern United States' ''

Cornus florida ''Cornus florida'', the flowering dogwood, is a species of flowering plant, flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to n ...

'' (flowering dogwood) is similar in appearance and possibly in chemical composition. ''

Cornus canadensis ''Cornus canadensis'' is a species of flowering plant in the dogwood family Cornaceae, native plant, native to eastern Asia and North America. Common names include Canadian dwarf cornel, Canadian bunchberry, quatre-temps, crackerberry, and creepi ...

'' has similar blossoms but grows as a

groundcover Groundcover or ground cover is any plant that grows low over an area of ground, which protects the topsoil from erosion and drought. In a terrestrial ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as the ...

Etymology

In 1806,

Meriwether Lewis Meriwether Lewis (August 18, 1774 – October 11, 1809) was an American explorer, soldier, politician, and public administrator, best known for his role as the leader of the Lewis and Clark Expedition, also known as the Corps of Discovery, with ...

noted that the species is similar in appearance to ''C. florida''. However, when Scottish botanist David Douglas encountered ''C. nuttalli'' on his expedition to the Pacific Northwest in the 1820s, he mistook it for ''C. florida'' and did not send seeds back to England. English botanist

Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an English botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle in the West Riding of Yorkshire a ...

was the first to describe the species for science while staying at

Fort Vancouver Fort Vancouver was a 19th-century fur trading post built in the winter of 1824–1825. It was the headquarters of the Hudson's Bay Company's Columbia Department, located in the Pacific Northwest. Named for Captain George Vancouver, the fort was ...

in the autumn of 1834. It was named ''nuttallii'' after him by his friend

John James Audubon John James Audubon (born Jean-Jacques Rabin, April 26, 1785 – January 27, 1851) was a French-American Autodidacticism, self-trained artist, natural history, naturalist, and ornithology, ornithologist. His combined interests in art and ornitho ...

. The common names comes from that of ''

Cornus sanguinea ''Cornus sanguinea'', the common dogwood or bloody dogwood, is a species of dogwood native to most of Europe and western Asia, from England and central Scotland east to the Caspian Sea. It is widely grown as an ornamental plant. Description It ...

'', the hard wood of which Northern Europeans used to make nails ("dags") during the medieval era. British Columbia Dogwood

Distribution and habitat

It occurs from the lowlands of southern

British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...

to the mountains of southern

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

. There exists an inland population in central

Idaho Idaho ( ) is a landlocked U.S. state, state in the Pacific Northwest and Mountain states, Mountain West subregions of the Western United States. It borders Montana and Wyoming to the east, Nevada and Utah to the south, and Washington (state), ...

, where it is considered critically imperiled. It occurs predominantly below in elevation. Cultivated examples are found as far north as

Haida Gwaii Haida Gwaii (; / , literally "Islands of the Haida people"), previously known as the Queen Charlotte Islands, is an archipelago located between off the British Columbia Coast, northern Pacific coast in the Canadian province of British Columbia ...

. It has high flood tolerance, and is common along streams with moist but well-drained soils, often on gentle slopes. Soil composition can range from clay to sandy loam, and it prefers a high humus content, moderate to high nutrient levels, and acidic soils with a pH from 5.5 to 6. It has low frost tolerance, and is usually found in low-elevation temperate or mesothermal climates. It is hardy to USDA zone 7.

Ecology

Value to animals

New sprouts are good

browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. In context of humans, it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing open sh ...

for both wild and domesticated

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

s, especially after a recent fire, but the mature foliage is usually ignored by all species except

slugs Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a sma ...

. The fruit are eaten by

deer mice ''Peromyscus'' is a genus of rodents. They are commonly referred to as deer mice or deermice, not to be confused with the chevrotain or "mouse deer". They are New World mice only distantly related to the common house and laboratory mouse, ''M ...

, pileated woodpeckers, the band-tailed pigeon, and bears. It provides habitat and cover to small birds such as Wilson's warbler, and small mammals including the red tree vole.

Diseases

Like the related ''Cornus florida'', it is very susceptible to dogwood

anthracnose A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...

, a

disease A disease is a particular abnormal condition that adversely affects the structure or function (biology), function of all or part of an organism and is not immediately due to any external injury. Diseases are often known to be medical condi ...

caused by the fungus '' Discula destructiva''. Fungal activity is greatest from May to July, although it can be active any time conditions are moist and the plant is growing. Infected leaves become blotched and drop, and defoliation can be extreme. Twigs and leaf buds are also impacted. This has killed many of the larger plants in the wild and has also restricted its use as an ornamental tree, to the point where it is considered threatening to the species in its native range.

Successional status

It is present in all stages of both primary and secondary

succession Succession is the act or process of following in order or sequence. Governance and politics *Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of ...

from new colonization on glacial outwash or areas destroyed by the

1980 eruption of Mount St. Helens In March 1980, a series of volcanic explosions and pyroclastic flows began at Mount St. Helens in Skamania County, Washington, United States. A series of Phreatic eruption, phreatic blasts occurred from the summit and escalated until a major ...

, to late seral and even climax communities. It is

shade tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in nursery (ho ...

but prefers sunlight in sufficiently humid conditions. It is adapted to a wide variety of

fire regime A fire regime is the pattern, frequency, and intensity of the bushfires and wildfires that prevail in an area over long periods of time. It is an integral part of fire ecology, and renewal for certain types of ecosystems. A fire regime describes th ...

s, with intervals ranging from just one year on dry sites, to 500 years or more in moist, riparian zones. The tree can survive low severity wildfires which are not hot enough to kill buds protected by bark. After being severely burned, ''Cornus nuttali'' typically resprouts from the root crownhowever, the resulting shoots are so palatable to

mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...

that they are at risk of being killed by over-browsing.

Uses

Some

Plateau Indian Indigenous peoples of the Northwest Plateau, also referred to by the phrase Indigenous peoples of the Plateau, and historically called the Plateau Indians (though comprising many groups) are Indigenous peoples of the Interior of British Columbi ...

tribes such as the

Nlaka'pamux The Nlakaʼpamux or Nlakapamuk ( ; ), also previously known as the ''Thompson'', '' Thompson River Salish'', ''Thompson Salish'', ''Thompson River Indians'' or ''Thompson River people'', and historically as the ''Klackarpun'', ''Haukamaugh'', ''K ...

used the bark as a brown dye. Those groups also used the bark for medicinal purposes as a blood purifier, lung strengthener, stomach treatment,

laxative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...

, and

emetic Vomiting (also known as emesis, puking and throwing up) is the forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenteritis, preg ...

. It is mostly prized as a cultivated ornamental.

In culture

It has been the provincial flower of

since 1956. It was once protected by law in the province (in an act which also protected '' Rhododendron macrophyllum'' and '' Trillium ovatum''), but this was repealed in 2002.

References

External links

Jepson Flora Project: ''Cornus nuttallii''

Plants of British Columbia: ''Cornus nuttallii''

Interactive Distribution Map of ''Cornus nuttallii''
* {{Taxonbar, from=Q220777 nuttallii Plants described in 1840 Trees of Northern America Ornamental trees Provincial symbols of British Columbia Trees of mild maritime climate Flora without expected TNC conservation status