''Pachydactylus'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
insectivorous
A robber fly eating a hoverfly
An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects.
The first vertebrate insectivores were ...
gecko
Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from .
Geckos ar ...
s,
lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia al ...
s in the
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Gekkonidae
Gekkonidae (the common geckos) is the largest family of geckos, containing over 950 described species in 64 genera. Members of the Gekkonidae comprise many of the most widespread gecko species, including house geckos ('' Hemidactylus''), tokay ...
. The genus is
endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to
Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and member
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
are commonly known as thick-toed geckos. The genus also displays rich
speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
, having 57 distinct species identified when compared to other closely related gecko genera like ''
Rhoptropus,'' most of which have emerged since 35
Ma.
It has been suggested that the reason for this rich speciation not from
adaptive radiation
In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
nor
nonadaptive radiation Nonadaptive radiations are a subset of evolutionary radiations (or species flocks) that are characterized by diversification that is not driven by resource partitioning. The species that are a part of a nonadaptive radiation will tend to have very ...
, but that the genus represents a clade somewhere between the two drivers of speciation.
''P.
bibronii'' geckos have been used by NASA as animal models for experimentation.
Description
The genus ''Pachydactylus'' is characterized by dilated toe tips, usually with undivided scansors. Body scales are small, granular and non-overlapping, with scattered, large keeled
tubercles
In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal.
In plants
A tubercle is generally a wart-like projection, ...
.
Coloration of Pachydactylus species varies, but is generally drab in color.
Presence of adhesive toe pads varies by species and habitat, with rock dwelling species of ''Pachydactylus'' retaining adhesive pads, but unambiguous independent loss of toe pads in sand dwelling and burrowing species like
''P. rangei''.
Body size in ''Pachydactylus'' varies across the 57 species, ranging from 35 to 115mm
Snout-Vent Length (SVL) with the
ancestral condition of a larger body size with adhesive toe pads to suit a generalist habitat.
Behavior
All observed
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of ''Pachydactylus'' are strictly
nocturnal
Nocturnality is an ethology, animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite.
Nocturnal creatures generally have ...
.
Habitat
''Pachydactylus'' species live in a diverse range of
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s across Southern Africa. Habitat varies by species, with some species preferring generalist habitats, human dwellings, rock-dwellings, and sand dwellings. Habitat preference typically varies by body size and retention of toe pads, which varies across the genus.
The body size of ''Pachydactylus'' geckos has been shown to correlate with their habitat range.
Diet
Lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia al ...
s of the genus ''Pachydactylus'' feed mainly on
arthropods
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, oft ...
, but have been observed eating small vertebrates.
Geographic range
The geographic range of the genus ''Pachydactylus'' is centered on
Southern Africa
Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number ...
, with some species reaching
East Africa
East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa:
Due to the histori ...
, the northernmost limit of their distribution. In South Africa's rugged
Richtersveld
The Richtersveld is a desert landscape characterised by rugged kloofs and high mountains, situated in the north-western corner of South Africa’s Northern Cape province. It is full of changing scenery from flat, sandy, coastal plains, to cragg ...
region, ''Pachydactylus'' geckos comprised 13 of 18 all gecko species surveyed.
Species
There are 57
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
that are recognized as being valid:
["''Pachydactylus'' ". The Reptile Database. www.reptile-database.org.]
*''
Pachydactylus acuminatus''
*''
Pachydactylus affinis'' –
Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name Transvaal.
* South African Republic (1856–1902; af, ...
thick-toed gecko
*''
Pachydactylus amoenus
''Pachydactylus amoenus'', also known as Namaqua banded gecko or Kamaggas gecko, is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well a ...
''
*''
Pachydactylus angolensis'' - Angola large-scaled gecko, Angolan thick-toed gecko
*''
Pachydactylus atorquatus''
*''
Pachydactylus austeni'' –
Austen's thick-toed gecko
*''
Pachydactylus barnardi
''Pachydactylus barnardi'', also known commonly as Barnard's rough gecko or Barnard's thick-toed gecko, is a species of lizard in the family Gekkonidae. The species is indigenous to Southern Africa.
Etymology
The specific name, ''barnardi'', ...
'' –
Barnard's thick-toed gecko
*''
Pachydactylus bicolor
The two-coloured thick-toed gecko (''Pachydactylus bicolor'') is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic isla ...
'' – velvety thick-toed gecko
*''
Pachydactylus boehmei''
*''
Pachydactylus capensis'' –
Cape
A cape is a clothing accessory or a sleeveless outer garment which drapes the wearer's back, arms, and chest, and connects at the neck.
History
Capes were common in medieval Europe, especially when combined with a hood in the chaperon. T ...
thick-toed gecko
*''
Pachydactylus caraculicus
The Angola banded thick-toed gecko (''Pachydactylus caraculicus'') is a species of lizard in the family Gekkonidae. It is found in southwestern Angola and northern Namibia
Namibia (, ), officially the Republic of Namibia, is a country in ...
'' –
Angola
, national_anthem = "Angola Avante"()
, image_map =
, map_caption =
, capital = Luanda
, religion =
, religion_year = 2020
, religion_ref =
, coordina ...
banded thick-toed gecko
*''
Pachydactylus carinatus
The Richtersveld gecko (''Pachydactylus carinatus'') is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains ...
''
*''
Pachydactylus etultra''
*''
Pachydactylus fasciatus
''Pachydactylus fasciatus'', also known as the striped thick-toed gecko, banded thick-toed gecko, thick-toed banded gecko, or Damaraland banded gecko, is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,0 ...
'' – banded thick-toed gecko
*''
Pachydactylus formosus
''Pachydactylus formosus'', also known as Smith's thick-toed gecko, southern rough gecko or Karoo gecko, is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents exce ...
'' –
Smith's thick-toed gecko
*''
Pachydactylus gaiasensis'' – Brandberg thick-toed gecko
*''
Pachydactylus geitje'' – ocellated thick-toed gecko
*''
Pachydactylus griffini''
*''
Pachydactylus haackei'' – Haacke's thick-toed gecko
*''
Pachydactylus katanganus''
*''
Pachydactylus kladaroderma'' – thin-skinned thick-toed gecko
*''
Pachydactylus kobosensis''
*''
Pachydactylus kochii
The Cape Cross thick-toed gecko (''Pachydactylus kochii''), also known commonly as Koch's gecko and Koch's thick-toed gecko, is a species of gecko, a lizard in the family Gekkonidae. The species is endemic to southern Africa.
Etymology
The spe ...
''
*''
Pachydactylus labialis'' –
Calvinia
Calvinia is a regional town in the Northern Cape province of South Africa named after the French religious reformer Jean Calvin. The town falls under the Hantam Local Municipality which forms part of the Namakwa District Municipality. The Calvinia ...
thick-toed gecko
*''
Pachydactylus latirostris'' – quartz gecko
*''
Pachydactylus macrolepis
The large-scaled banded gecko (''Pachydactylus macrolepis'') is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic islan ...
'' – large-scaled banded gecko
*''
Pachydactylus maculatus'' – spotted thick-toed gecko
*''
Pachydactylus maraisi
''Pachydactylus maraisi'' is a species of lizard in the family Gekkonidae. The species is endemic to Namibia.
Etymology
The specific name, ''maraisi'' is in honor of African herpetologist Johan Marais
Johannes Hendricus Marais (born 2 ...
''
*''
Pachydactylus mariquensis
''Pachydactylus mariquensis'', also known as the common banded gecko, Ceres thick-toed gecko, or Marico thick-toed gecko, is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across a ...
'' – Marico thick-toed gecko
*''
Pachydactylus mclachlani
''Pachydactylus mclachlani'' is a species of lizard in the family Gekkonidae. The species is endemic to Namibia.
Etymology
The specific name, ''mclachlani'', is in honor of Geoffrey Roy McLachlan (1923–2005), who was a South African orni ...
''
*''
Pachydactylus monicae''
*''
Pachydactylus montanus''
*''
Pachydactylus namaquensis'' – Namaqua thick-toed gecko
*''
Pachydactylus oculatus'' – golden spotted thick-toed gecko
*''
Pachydactylus oreophilus'' –
Kaokoveld
The Kaokoveld Desert is a coastal desert of northern Namibia and southern Angola.
Setting
The Kaokoveld Desert occupies a coastal strip covering , from 13° to 21°S and is bounded by the Atlantic Ocean to the west, the Namibian savanna woodla ...
thick-toed gecko
*''
Pachydactylus oshaughnessyi
''Pachydactylus oshaughnessyi'' is a species of lizard in the family Gekkonidae. The species is endemic to southern Africa.
Etymology
The specific name, ''oshaughnessyi'', is in honor of Arthur O'Shaughnessy, who was a British herpetologist an ...
''
*''
Pachydactylus otaviensis''
*''
Pachydactylus parascutatus''
*''
Pachydactylus punctatus
''Pachydactylus punctatus'', also known as the pointed thick-toed gecko or speckled thick-toed gecko, is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except ...
'' – speckled thick-toed gecko
*''
Pachydactylus purcelli''
*''
Pachydactylus rangei'' –
Namib
The Namib ( ; pt, Namibe) is a coastal desert in Southern Africa. The name is of Khoekhoegowab origin and means "vast place". According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Nami ...
sand gecko
*''
Pachydactylus reconditus
''Pachydactylus reconditus'' is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyl ...
''
*''
Pachydactylus robertsi
''Pachydactylus robertsi'', commonly known as the large-scaled gecko, shielded thick-toed gecko, or ''Rauhschuppen-Dickfingergecko'' in German, is an African species of gecko.
Geographic range
''P. robertsi'' is found near Keetmanshoop and Kara ...
''
*''
Pachydactylus rugosus'' – rough thick-toed gecko
*''
Pachydactylus sansteynae'' – San Steyn's thick-toed gecko
*''
Pachydactylus scherzi'' – Sherz's thick-toed gecko
*''
Pachydactylus scutatus
''Pachydactylus scutatus'', also known as the large-scaled gecko or shielded thick-toed gecko, is a species of lizard in the family Gekkonidae. It is found in Namibia, South Africa, and Angola
, national_anthem = " Angola Avante"( ...
'' – large-scaled thick-toed gecko
*''
Pachydactylus serval
The western spotted thick-toed gecko (''Pachydactylus serval'') is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic is ...
'' – western spotted thick-toed gecko
*''
Pachydactylus tigrinus'' – tiger thick-toed gecko
*''
Pachydactylus tsodiloensis
The Tsodilo thick-toed gecko (''Pachydactylus tsodiloensis'') is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic isla ...
'' –
Tsodilo thick-toed gecko
*''
Pachydactylus vansoni'' – Van Son's thick-toed gecko
*''
Pachydactylus vanzyli''
*''
Pachydactylus visseri
Visser's gecko (''Pachydactylus visseri'') is a species of lizard in the family Gekkonidae. The species is native to southern Africa.
Etymology
The specific name, ''visseri'', is in honor of South African herpetologist John Duckitt Visser ( ...
''
*''
Pachydactylus wahlbergii''
*''
Pachydactylus waterbergensis
''Pachydactylus waterbergensis'' is a species of lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is para ...
''
*''
Pachydactylus weberi
Weber's thick-toed gecko (''Pachydactylus weberi'') is a species of lizard in the family Gekkonidae. The species is native to southern Africa.
Etymology
The specific name, ''weberi'', is in honor of German-Dutch zoologist Max Wilhelm Carl Webe ...
'' –
Weber's thick-toed gecko
*''
Pachydactylus werneri
Werner's thick-toed gecko (''Pachydactylus werneri'') is a species of lizard in the family Gekkonidae. The species is endemic to Namibia.
Etymology
The specific name, ''werneri'', is in honor of Austrian herpetologist Franz Werner. Beolens, ...
''
''
Nota bene
(, or ; plural form ) is a Latin phrase meaning "note well".
It is often abbreviated as NB, n.b., or with the ligature
and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atte ...
'': A
binomial authority
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Pachydactylus''.
References
Further reading
*
*
*
*
External links
*
{{Taxonbar, from=Q2482361
Geckos of Africa
Lizard genera
Taxa named by Arend Friedrich August Wiegmann