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''Pachydactylus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...
s,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Gekkonidae Gekkonidae (the common geckos) is the largest family of geckos, containing over 950 described species in 62 genera. The Gekkonidae contain many of the most widespread gecko species, including house geckos (''Hemidactylus''), the tokay gecko (''Ge ...
. The genus is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, and member
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are commonly known as thick-toed geckos. The genus also displays rich
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
, having 58 distinct species identified when compared to other closely related gecko genera like '' Rhoptropus,'' most of which have emerged since 35 Ma. It has been suggested that the reason for this rich speciation not from
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
nor nonadaptive radiation, but that the genus represents a clade somewhere between the two drivers of speciation. ''P. bibronii'' geckos have been used by NASA as animal models for experimentation.


Description of the ''Pachydactylus'' genus

The genus ''Pachydactylus'' is characterized by dilated toe tips, usually with undivided scansors. Body scales are small, granular and non-overlapping, with scattered, large keeled tubercles. Coloration of Pachydactylus species varies, but is generally drab in color. Presence of adhesive toe pads varies by species and habitat, with rock dwelling species of ''Pachydactylus'' retaining adhesive pads, but unambiguous independent loss of toe pads in sand dwelling and burrowing species like ''P. rangei''. Body size in ''Pachydactylus'' varies across the 58 species, ranging from 35 to 115mm Snout-Vent Length (SVL) with the ancestral condition of a larger body size with adhesive toe pads to suit a generalist habitat.


Behavior

All observed
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of ''Pachydactylus'' are strictly
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
.
Wikibooks Wikibooks (previously called ''Wikimedia Free Textbook Project'' and ''Wikimedia-Textbooks'') is a wiki-based Wikimedia project hosted by the Wikimedia Foundation for the creation of free content digital textbooks and annotated texts that anyon ...


Habitat of ''Pachydactylus'' geckos

''Pachydactylus'' species live in a diverse range of
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s across Southern Africa. Habitat varies by species, with some species preferring generalist habitats, human dwellings, rock-dwellings, and sand dwellings. Habitat preference typically varies by body size and retention of toe pads, which varies across the genus. The body size of ''Pachydactylus'' geckos has been shown to correlate with their habitat range.


Diet

Lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s of the genus ''Pachydactylus'' feed mainly on
arthropods Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
, but have been observed eating small vertebrates.


Geographic range

The geographic range of the genus ''Pachydactylus'' is centered on
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, with some species reaching
East Africa East Africa, also known as Eastern Africa or the East of Africa, is a region at the eastern edge of the Africa, African continent, distinguished by its unique geographical, historical, and cultural landscape. Defined in varying scopes, the regi ...
, the northernmost limit of their distribution. In South Africa's rugged
Richtersveld The Richtersveld is a desert landscape characterised by rugged kloofs and high mountains, situated in the north-western corner of South Africa’s Northern Cape province. It is full of changing scenery from flat, sandy, coastal plains, to cragg ...
region, ''Pachydactylus'' geckos comprised 13 of 18 all gecko species surveyed.


Species

There are 58
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
that are recognized as being valid: www.reptile-database.org. *'' Pachydactylus acuminatus'' *'' Pachydactylus affinis'' –
Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name ''Transvaal''. * South African Republic (1856–1902; ...
thick-toed gecko *'' Pachydactylus amoenus'' *'' Pachydactylus angolensis'' – Angola large-scaled gecko, Angolan thick-toed gecko *'' Pachydactylus atorquatus'' *'' Pachydactylus austeni'' – Austen's thick-toed gecko *'' Pachydactylus barnardi'' – Barnard's thick-toed gecko xiii + 296 pp. (''Pachydactylus barnardi'', p. 17; ''P. weberi'', p. 280). *'' Pachydactylus bicolor'' – velvety thick-toed gecko *'' Pachydactylus boehmei'' *'' Pachydactylus capensis'' –
Cape A cape is a clothing accessory or a sleeveless outer garment of any length that hangs loosely and connects either at the neck or shoulders. They usually cover the back, shoulders, and arms. They come in a variety of styles and have been used th ...
thick-toed gecko *'' Pachydactylus caraculicus'' –
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
banded thick-toed gecko *'' Pachydactylus carinatus'' *'' Pachydactylus etultra'' *'' Pachydactylus fasciatus'' – banded thick-toed gecko *'' Pachydactylus formosus'' – Smith's thick-toed gecko *'' Pachydactylus gaiasensis'' – Brandberg thick-toed gecko *'' Pachydactylus geitje'' – ocellated thick-toed gecko *'' Pachydactylus griffini'' *'' Pachydactylus haackei'' – Haacke's thick-toed gecko *'' Pachydactylus katanganus'' *'' Pachydactylus kladaroderma'' – thin-skinned thick-toed gecko *'' Pachydactylus kobosensis'' *'' Pachydactylus kochii'' *'' Pachydactylus labialis'' –
Calvinia Calvinia is a regional town in the Northern Cape province of South Africa named after the French people, French religious reformer Jean Calvin. The town falls under the Hantam Local Municipality which forms part of the Namakwa District Municipalit ...
thick-toed gecko *'' Pachydactylus latirostris'' – quartz gecko *'' Pachydactylus macrolepis'' – large-scaled banded gecko *'' Pachydactylus maculatus'' – spotted thick-toed gecko *'' Pachydactylus maiatoi'' – Maiato's thick-toed gecko *'' Pachydactylus maraisi'' *'' Pachydactylus mariquensis'' – Marico thick-toed gecko *'' Pachydactylus mclachlani'' *'' Pachydactylus monicae'' *'' Pachydactylus montanus'' *'' Pachydactylus namaquensis'' – Namaqua thick-toed gecko *'' Pachydactylus oculatus'' – golden spotted thick-toed gecko *'' Pachydactylus oreophilus'' – Kaokoveld thick-toed gecko *'' Pachydactylus oshaughnessyi'' *'' Pachydactylus otaviensis'' *'' Pachydactylus parascutatus'' *'' Pachydactylus punctatus'' – speckled thick-toed gecko *'' Pachydactylus purcelli'' *'' Pachydactylus rangei'' –
Namib The Namib ( ; ) is a coastal desert in Southern Africa. According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Namibia, and northwest South Africa, extending southward from the Carunjamba Ri ...
sand gecko *'' Pachydactylus reconditus'' *'' Pachydactylus robertsi'' *'' Pachydactylus rugosus'' – rough thick-toed gecko *'' Pachydactylus sansteynae'' – San Steyn's thick-toed gecko *'' Pachydactylus scherzi'' – Sherz's thick-toed gecko *'' Pachydactylus scutatus'' – large-scaled thick-toed gecko *'' Pachydactylus serval'' – western spotted thick-toed gecko *'' Pachydactylus tigrinus'' – tiger thick-toed gecko *'' Pachydactylus tsodiloensis'' –
Tsodilo The Tsodilo Hills () are a UNESCO World Heritage Site (WHS), consisting of rock art, rock shelters, depressions, and caves in Botswana, Southern Africa. It gained its WHS listing in 2001 because of its unique religious and spiritual significance ...
thick-toed gecko *'' Pachydactylus vansoni'' – Van Son's thick-toed gecko *'' Pachydactylus vanzyli'' *'' Pachydactylus visseri'' *'' Pachydactylus wahlbergii'' *'' Pachydactylus waterbergensis'' *'' Pachydactylus weberi'' – Weber's thick-toed gecko *'' Pachydactylus werneri'' ''
Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': A binomial authority in parentheses indicates that the species was originally described in a genus other than ''Pachydactylus''.


References


Further reading

* * xii + 436 pp. + Plates I–XXXII. (Genus ''Pachydactylus'', p. 200). * (Third Revised edition, Second impression). 399 pp. (Genus ''Pachydactylus'', pp. 249–250). * (''Pachydactylus'', new genus, p. 19). (in Latin).


External links

* {{Authority control Geckos of Africa Lizard genera Taxa named by Arend Friedrich August Wiegmann