''Pinus orizabensis'', the Orizaba pinyon, is a

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...

in the

pinyon pine The pinyon or piñon pine group grows in southwestern North America, especially in New Mexico, Colorado, Arizona, and Utah, with the single-leaf pinyon pine just reaching into southern Idaho. The trees yield edible Pine nut, nuts, which are a sta ...

group,

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to central

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

. It is considered also as a sub-species of ''

Pinus cembroides ''Pinus cembroides'', also known as pinyon pine, Mexican pinyon, Mexican nut pine, and Mexican stone pine, is a pine in the pinyon pine group. It is a small pine growing to about with a trunk diameter of up to . It is native to western North Am ...

'' which is classified as ''Pinus cembroides'' subsp. ''orizabensis'' D.K.Bailey. It has larger seeds than the type. The range is localised, confined to a small area in the eastern

Eje Volcánico Transversal The Trans-Mexican Volcanic Belt (), also known as the Transvolcanic Belt and locally as the (''Snowy Mountain Range''), is an active volcanic belt that covers central-southern Mexico. Several of its highest peaks have snow all year long, and du ...

range (Trans-Mexican Volcanic Belt), in the states of

Puebla Puebla, officially the Free and Sovereign State of Puebla, is one of the 31 states that, along with Mexico City, comprise the Federal Entities of Mexico. It is divided into 217 municipalities and its capital is Puebla City. Part of east-centr ...

and

Veracruz Veracruz, formally Veracruz de Ignacio de la Llave, officially the Free and Sovereign State of Veracruz de Ignacio de la Llave, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entit ...

. It occurs at high altitudes, mostly from , in a cooler, moister climate than the other pinyon pines.

Description

It is a small to medium-size

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

, reaching tall and with a trunk diameter of up to 50 cm. The bark is dark brown, thick and deeply fissured at the base of the trunk. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

('needles') are in mixed fascicles of three and four, slender, 3–6 cm long, and deep green to blue-green, with

stomata In botany, a stoma (: stomata, from Greek ''στόμα'', "mouth"), also called a stomate (: stomates), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange between the internal air spa ...

confined to a bright white band on the inner surfaces.

Cones

The

cones In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines, ...

are globose to ovoid, 4–7 cm long and 3–5 cm broad when closed, green at first, ripening yellow-brown when 16–18 months old, with only a small number of thin scales, typically 6-18 fertile scales. The cones open to 5–7 cm broad when mature, holding the

seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...

s on the scales after opening. The seeds are 12–15 mm long, with a thick shell, a pink

endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...

, and a vestigial 2 mm wing; they are dispersed by the

Mexican jay The Mexican jay (''Aphelocoma wollweberi'') Etymology: ''Aphelocoma'', from Latinized Ancient Greek ''apheles-'' (from ἀφελής-) "simple" + Latin ''coma'' (from Greek ''kome'' κόμη) "hair", in reference to the lack of striped or banded ...

, which plucks the seeds out of the open cones. The jay, which uses the seeds as a major food resource, stores many of the seeds for later use, and some of these stored seeds are not used and are able to grow into new trees.

History

''Pinus orizabensis'' is the most recent

to be described, discovered by

Dana K. Bailey Dana may refer to: Businesses and organisations * Dana (company), a Slovenian beverage company * Dana (payment service), in Indonesia * Dana Air, a Nigerian airline * Dana College, formerly in Nebraska, U.S. * Dana Energy, an Iranian oil and g ...

in 1983 when examining an unusual pinyon cultivated at the

Royal Botanic Gardens, Kew Royal Botanic Gardens, Kew is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. An internationally important botanical research and education institution, it employs 1,10 ...

; it was found to match wild specimens from the

Pico de Orizaba Citlaltépetl (from Nahuan languages, Náhuatl = star, and = mountain), otherwise known as Pico de Orizaba, is an active volcano, the highest mountain in Mexico and Table of the highest major summits of North America, third highest in North Ame ...

. At first it was described as a

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

Mexican pinyon ''Pinus cembroides'', also known as pinyon pine, Mexican pinyon, Mexican nut pine, and Mexican stone pine, is a pine in the pinyon pine group. It is a small pine growing to about with a trunk diameter of up to . It is native to western North Am ...

, but further research by Bailey & Hawksworth and others has shown that it is better treated as a distinct

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

. Some botanists still include it in Mexican pinyon though, despite their occurring together in some sites without evidence of hybridisation. Orizaba pinyon shows better adaptation to rainy temperate climate like

England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...

than ''Pinus cembroides'' which grows in dryer habitats. ''Pinus orizabensis'' is most closely allied to

Johann's pinyon ''Pinus johannis'', the Johann's pine, is a pine in the pinyon pine group, native to North America. The range extends from southeast Arizona and southwest New Mexico, United States, south in Mexico along the Sierra Madre Occidental and the Sie ...

and

Potosi pinyon ''Pinus culminicola'', commonly known as Potosí pinyon or Potosí Piñón, is a pine in the pinyon pine group, native and endemic to northeast Mexico. The range is highly localised, confined to a small area of high summits in the northern Sier ...

, with which it shares the leaf structure with the stomata confined to the inner faces; it differs from these in the larger cones and seeds, and from the latter in fewer needles per fascicle (3-4 vs 5). Like these two, the white-glaucous inner surfaces of the needles make it a very attractive small tree, suitable for

park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...

s and large

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...

s. The edible (

pine nut Pine nuts, also called piñón (), pinoli (), or pignoli, are the edible seeds of pines (family Pinaceae, genus ''Pinus''). According to the Food and Agriculture Organization, only 29 species provide edible nuts, while 20 are traded locall ...

) seeds are collected in Mexico to a small extent.

References

Photo of cones (''scroll half-way down'')
{{Taxonbar, from=Q3308263 orizabensis Endemic flora of Mexico Trees of Northern America Flora of the Trans-Mexican Volcanic Belt Edible nuts and seeds Bird food plants Trees of mild maritime climate Near threatened plants Taxobox binomials not recognized by IUCN