Opisthocomiformes
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Opisthocomidae is a family of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, the only named family within the order Opisthocomiformes. The only living representative is the
hoatzin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
(''Opisthocomus hoazin'') which lives in the
Amazon Amazon most often refers to: * Amazon River, in South America * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon (company), an American multinational technology company * Amazons, a tribe of female warriors in Greek myth ...
and the
Orinoco delta The Orinoco Delta is a vast river delta of the Orinoco River, located in eastern Venezuela. Location The Orinoco Delta is one of the eight natural regions of Venezuela. It covers the whole of Delta Amacuro State and a few square kilometers o ...
in South America. Several fossil species have been identified, including one from Africa and one from Europe.


Phylogeny

The phylogeny below is based on the work of Hughes & Baker 1999 and Mayr & De Pietri 2014. Traditionally classified among the fowl-like birds (
Galliformes Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
), recent studies have favored Opisthocomidae's placement within the
Neoaves Neoaves is a clade that consists of all modern bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard ...
.


Taxonomy

* Family Opisthocomidae Swainson 1837Mikko's Phylogeny Archiv

** Genus ?†'' Foro (bird), Foro'' Olson 1992 (mid-Eocene, USA) - cuculiform? *** Species †'' Foro panarium'' Olson 1992 ** Genus ?†'' Onychopteryx'' Cracraft 1971 (Early Eocene of Argentina) – falconid? A ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
'' *** Species †'' Onychopteryx simpsoni'' Cracraft 1971 ** Genus †'' Protoazin'' Mayr & De Pietri 2014 (late Eocene of France) *** Species †'' Protoazin parisiensis'' Mayr & De Pietri 2014 ** Genus †'' Namibiavis'' Mourer-Chauviré 2003 (Middle Miocene of Namibia) *** Species †'' Namibiavis senutae'' Mourer-Chauviré 2003 ** Genus †'' Hoazinavis'' Alvarenga, Mayr & Mourer-Chauviré 2011 (Late Oligocene and Early Miocene of Brazil) *** Species †'' Hoazinavis lacustris'' Alvarenga, Mayr & Mourer-Chauviré 2011 ** Genus †'' Hoazinoides'' Miller 1953 (middle Miocene of Colombia) *** Species †'' Hoazinoides magdalenae'' Miller 1953 ** Genus '' Opisthocomus'' Illiger 1811 *** Species ''
Opisthocomus hoazin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
'' (Müller 1776) Illiger 1811 'Phasianus hoazin'' Müller 1776


Description

The only living representative is the
hoatzin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
(''Opisthocomus hoazin''), which lives in the
Amazon Amazon most often refers to: * Amazon River, in South America * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon (company), an American multinational technology company * Amazons, a tribe of female warriors in Greek myth ...
and the
Orinoco delta The Orinoco Delta is a vast river delta of the Orinoco River, located in eastern Venezuela. Location The Orinoco Delta is one of the eight natural regions of Venezuela. It covers the whole of Delta Amacuro State and a few square kilometers o ...
in South America. Several fossil species have been identified, including one from Africa and one from Europe. The
hoatzin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
is a stunning, colourful bird from South America. The hoatzin is found in the rainforest of the Amazon. It has a long, permanently erected crest which gives the species a distinctive silhouette. This gregarious species is often found in groups of more than 40 birds, and up to 100 birds during breeding season which occurs during rainfalls. The nest is a flat platform in bush or tree above water. The chicks are semi-precocial and leave the nest at about two to three weeks of age. The adults feed them with semi-digested slimy mass from their crops, and the young are fed during the first four or five months of their life. The young nestlings can jump out into the water, and swim with wings and feet, in order to escape predators at nest. Hoatzin embryos are known to develop very quickly compared to other birds. Hoatzin (Opisthocomus hoazin) Rio Napo 2.jpg Hoatzin (Opisthocomus hoazin) head Rio Napo.jpg The
hoatzin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
lives in tropical forested wetlands of 200 to 500 meters elevation. The hoatzin is an
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
species and is folivorous, feeding on leaves, flowers and fruits of a small number of plant species. It has a large crop that uses bacteria to break down the leaves that it eats. The hoatzin is unique because it has bacteria in its crop that breaks down its food while it is still in the crop. This is a process called foregut fermentation. It rarely drinks because its diet is high in water. The hoatzin is sedentary and widespread in suitable habitat. They are noisy birds, calling in unison, uttering large numbers of varied sounds. In addition to foregut fermentation, the
hoatzin The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthoco ...
has a highly modified skeleton to accommodate its large crop, and in the young of this species, wing claws at the wrist joint which are used to climb among the branches of the nest tree.


References

{{Taxonbar, from=Q942816 Bird families Opisthocomiformes Herbivorous vertebrates Extant Eocene first appearances