Open-pollinated
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"Open pollination" and "open pollinated" refer to a variety of concepts in the context of the sexual reproduction of plants. Generally speaking, the term refers to plants pollinated naturally by birds, insects, wind, or human hands. Controlled pollination is the process of collecting the pollen variety from live flowers during the bloom season and processing them, and re-introducing the pollen back into the orchard via a backpack blower or dusting into the beehives. Increased yields can be accomplished and vary from 15% to 25% depending on application methods, timing, and weather conditions. A large variety of stone fruits are receptive to this process, i.e. almonds, avocados, cherries, olives, plums, etc. Controlled pollination is beneficial in times when bee flight either is hampered by bad weather or the lack of bees to pollinate enough orchards during the blooming season. Some growers do the application via aircraft and/or drones. This citation was added by Steve Urner, owner of Pollen Application Service. Studies are available. WWW.Pollenapplication .com


True-breeding definition

"Open pollinated" generally refers to seeds that will "breed true". When the plants of an open-pollinated variety self-pollinate, or are pollinated by another representative of the same variety, the resulting seeds will produce plants roughly identical to their parents. This is in contrast to the seeds produced by plants that are the result of a recent cross (such as, but not confined to, an
F1 hybrid F1 hybrid (also known as filial 1 hybrid) is the first filial generation of offspring of distinctly different parental types. F1 hybrids are used in genetics, and in selective breeding, where the term F1 crossbreed may be used. The term is somet ...
), which are likely to show a wide variety of differing characteristics. Open-pollinated varieties are also often referred to as standard varieties or, when the seeds have been saved across generations or across several decades,
heirloom In popular usage, an heirloom is something that has been passed down for generations through family members. Examples are a family bible, antiques, weapons or jewellery. The term originated with the historical principle of an heirloom in ...
varieties. While heirlooms are usually open-pollinated, open-pollinated seeds are not necessarily heirlooms; open-pollinated varieties are still being developed. One of the challenges in maintaining an open-pollinated variety is avoiding introduction of pollen from other strains. Based on how broadly the pollen for the plant tends to
disperse Disperse was a Christian rock band from Southern Indiana active from 1996 to 2004. The band was formerly known, with an adjusted roster, as "Stuff." Members (as of 2004 dispersion) :Chelsey Pieratt - Vocals :Chris Billings - Lead Guitar :And ...
, it can be controlled to varying degrees by
greenhouse A greenhouse is a structure that is designed to regulate the temperature and humidity of the environment inside. There are different types of greenhouses, but they all have large areas covered with transparent materials that let sunlight pass an ...
s, tall wall enclosures, field isolation, or other techniques. Because they breed true, the seeds of open-pollinated plants are often saved by home gardeners and farmers. Popular examples of open-pollinated plants include
heirloom tomato An heirloom tomato (also called heritage tomato in the UK) is an open-pollinated, non-hybrid heirloom cultivar of tomato. They are classified as family heirlooms, commercial heirlooms, mystery heirlooms, or created heirlooms. They usually have ...
es, beans, peas, and many other garden vegetables.


Uncontrolled pollination definition

A second use of the term "open pollination" refers to
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
by
insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...
,
birds Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
,
wind Wind is the natural movement of atmosphere of Earth, air or other gases relative to a planetary surface, planet's surface. Winds occur on a range of scales, from thunderstorm flows lasting tens of minutes, to local breezes generated by heatin ...
, or other
natural Nature is an inherent character or constitution, particularly of the ecosphere or the universe as a whole. In this general sense nature refers to the laws, elements and phenomena of the physical world, including life. Although humans are part ...
mechanisms. This can be contrasted with
cleistogamy Cleistogamy is a type of automatic self-pollination of certain plants that can propagate by using non-opening, self-pollinating flowers. Especially well known in peanuts, peas, and pansies, this behavior is most widespread in the grass family. ...
, closed pollination, which is one of the many types of
self pollination Self-pollination is a form of pollination in which pollen arrives at the stigma of a flower (in flowering plants) or at the ovule (in gymnosperms) of the same plant. The term cross-pollination is used for the opposite case, where pollen from one ...
.Kearns, C.A.; Inouye, D.W. 1993. Techniques for pollination biologists. University Press of Colorado, Niwot, CO. When used in this sense, open pollination may contrast with controlled pollination, a procedure used to ensure that all seeds of a crop are descended from parents with known traits, and are therefore more likely to have the desired traits. The
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s of open-pollinated
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s will produce new
generation A generation is all of the people born and living at about the same time, regarded collectively. It also is "the average period, generally considered to be about 20–⁠30 years, during which children are born and grow up, become adults, and b ...
s of those plants; however, because breeding is uncontrolled and the pollen (male parent) source is unknown, open pollination may result in plants that vary widely in genetic traits. Open pollination may increase
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
. Some plants (such as many crops) are primarily self pollenizing and also breed true, so that even under open pollination conditions the next generation will be (almost) the same. Even among true breeding organisms, some variation due to
genetic recombination Genetic recombination (also known as genetic reshuffling) is the exchange of genetic material between different organisms which leads to production of offspring with combinations of traits that differ from those found in either parent. In eukaryot ...
or to mutation can produce a few "off types".


Relationship to hybridization

Hybrid pollination, a type of controlled pollination in which the pollen comes from a different strain (or
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
), can be used to increase crop suitability, especially through
heterosis Heterosis, hybrid vigor, or outbreeding enhancement is the improved or increased function of any biological quality in a hybrid offspring. An offspring is heterotic if its traits are enhanced as a result of mixing the genetic contributions o ...
. The resulting hybrid strain can sometimes be
inbred Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders an ...
and selected for desired traits until a strain that breeds true by open pollination is achieved. The result is referred to as an inbred hybrid strain. To add some confusion, the term hybrid inbred applies to hybrids that are made from selected inbred lines that have certain desired characteristics (see
inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
). The latter type of hybrid is sometimes designated
F1 hybrid F1 hybrid (also known as filial 1 hybrid) is the first filial generation of offspring of distinctly different parental types. F1 hybrids are used in genetics, and in selective breeding, where the term F1 crossbreed may be used. The term is somet ...
, i.e. the first hybrid (filial) generation whose parents were (different) inbred lines.


References


Further reading

*{{cite web , url= http://www.garden.org/subchannels/care/seeds?q=show&id=293 , title= Hybrid or Open Pollinated , access-date= 2008-03-17 , author= Ben Watson , work= Gardening Articles: Care :: Seeds & Propagation , publisher= National Gardening Association , pages= 1–6 Pollination