Olenelloidea
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The Olenelloidea are a superfamily of
trilobites Trilobites (; meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, ...
, a group of
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
marine
arthropods Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
. They lived during the late
Lower Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovici ...
and species occurred on all paleocontinents.


Taxonomy

Palmer and Repina assigned '' Gabriellus'' and '' Callavia'' to the Olenelloidea, but the ocular lobes in these two genera circumscribe the entire lateral border of the frontal lobe of the glabella, and thus, do not belong to this superfamily, but to the Judomioidea. Geyer assigned '' Cambropallas'' to the Olenelloidea, but it lacks the diagnostic features and appears to be closely related to '' Andalusiana'', and is now considered to be an advanced member of the Nevadioidea.


Distribution

In western Laurentia Fallotaspididae and Archaeaspididae proceed the first Olenelloidea. The Holmiidae occur at the late Atdabanian, and are contemporary with the Nevadioidea. They are followed by Biceratopsidae and Olenellidae.


Description

As with most early trilobites, the Olenelloidea have an almost flat exoskeleton, that is only thinly calcified, and has crescent-shaped eye ridges. As part of the Olenellina suborder, the Olenelloidea lack dorsal sutures. The superfamily can be distinguished from all other
Olenellina Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. Olenellina are arguably the earliest trilobites in the fossil record as members ...
by features of the
cephalon Cephalon, Inc. was an American biopharmaceutical company co-founded in 1987 by pharmacologist Frank Baldino Jr., Frank Baldino, Jr., neuroscientist Michael Lewis, and organic chemist James C. Kauer—all three former scientists with the DuPont ...
and in particular the
glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior ...
. The backward portion of the third lateral lobe of the glabella counted from the back (L3) bulges laterally relative to the first lateral lobe (L1). The front end of the eye lobe (or ocular lobe) is connected only to the back half of the frontal lobe (AL or L4) of the glabella, and merges smoothly with it (rather than merely contacting it). The ocular lobe consists of an inner and outer band.


References

Olenellina Cambrian trilobites Cambrian first appearances Cambrian Series 2 extinctions Arthropod superfamilies {{Redlichiida-stub