''Notosuchus'' (; 'southern crocodile') is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of South American

notosuchia Notosuchia is a clade of primarily Gondwanan mesoeucrocodylian Crocodylomorpha, crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogeny, phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group ...

crocodyliform Crocodyliformes is a clade of Crurotarsi, crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the ...

s. It was terrestrial, living approximately 85 million years ago in the

Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...

stage of the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

Description

''Notosuchus'' was relatively small, reaching in length and a weight of . Remains have been found in the

Bajo de la Carpa Formation The Bajo de la Carpa Formation is a geologic formation of the Neuquén Basin that outcrop, crops out in northern Patagonia, in the provinces of Río Negro Province, Río Negro and Neuquén Province, Neuquén, Argentina. It is the oldest of two for ...

Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

. First named in 1896, ''Notosuchus'' was the first known notosuchian. The type species is ''N. terrestris''. A second species, ''N. lepidus'', was named in 1957. A paper published in 2008 by Fiorelli and Calvo described new remains of the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

''N. terrestris''. In it, the authors suggested that the skull would have supported a short trunk, or "hog's snout" as well as fleshy upper and lower lips. The anteriorly directed nares and the absence of a bony

nasal septum The nasal septum () separates the left and right airways of the Human nose, nasal cavity, dividing the two nostrils. It is Depression (kinesiology), depressed by the depressor septi nasi muscle. Structure The fleshy external end of the nasal s ...

(which presumably indicates cartilaginous tissue serving its place) provide evidence for a trunk-like snout, while striations on the surface of the

nasal bone The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Eac ...

s and the lower jaw most likely acted as attachment points for the nasolabial muscles and the depressor muscle, respectively.Fiorelli, L. E. (2005). "Nuevos restos de ''Notosuchus terrestris'' Woodward, 1896 (Crocodyliformes: Mesoeucrocodylia) del Cretácico Superior (Santoniano) de la Provincia de Neuquén, Patagonia, Argentina". 79p. Tesis (Grado) Universidad Nacional de Córdoba, Córdoba. Additionally, the authors suggested that a lateral rim on the

dentary In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone ...

as well as numerous aligned neurovascular foramina are evidence of soft

cheek The cheeks () constitute the area of the face below the eyes and between the nose and the left or right ear. ''Buccal'' means relating to the cheek. In humans, the region is innervated by the buccal nerve. The area between the inside of th ...

-like muscular tissue. The function of the trunk was likely used for searching for food by sniffing the ground in a manner similar to extant suids and

peccaries Peccaries (also javelinas or skunk pigs) are pig-like ungulates of the family Tayassuidae (New World pigs). They are found throughout Central and South America, Trinidad in the Caribbean, and in the southwestern area of North America. Peccari ...

, while the cheeks would aid in

mastication Chewing or mastication is the process by which food is comminution, crushed and ground by the teeth. It is the first step in the process of digestion, allowing a greater surface area for digestive enzymes to break down the foods. During the mast ...

by preventing food loss. A re-description of the skull material has since lent evidence toward minimal soft-tissue enhancement to the snout of ''Notosuchus''.

Diet

References

{{Authority control Notosuchia Late Cretaceous crocodylomorphs of South America Terrestrial crocodylomorphs Bajo de la Carpa Formation Fossil taxa described in 1896 Taxa named by Arthur Smith Woodward Prehistoric pseudosuchian genera Cretaceous Argentina