Notophyll
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Leaf size of plants can be described using the terms ''megaphyll'', ''macrophyll'', ''mesophyll'', ''microphyll'', ''nanophyll'' and ''leptophyll'' (in descending order) in a classification devised in 1934 by
Christen C. Raunkiær Christen Christensen Raunkiær (29 March 1860 – 11 March 1938) was a Danish botanist, who was a pioneer of plant ecology. He is mainly remembered for his scheme of plant strategies to survive an unfavourable season ("life forms") and his demon ...
and since modified by others. Definitions vary, some referring to length and others to area. Raunkiaer's original definitions were by leaf area, and differed by a factor of nine at each stage. Some authors simplified the system to make it specific to particular climates, and have introduced extra terms including ''notophyll'', ''picophyll'', ''platyphyll'' and ''subleptophyll''. In
ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
, microphyll and similar terms based on blade size of the leaf are used to describe a
flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...
, for example, a "microphyll rainforest" is often defined as a forest where the dominant trees have leaves less than 7.5 cm in length.


Raunkiaer's work

Christen C. Raunkiaer proposed using leaf size as a relatively easy measurement that could be used to compare the adaptation of a plant community to dryness.
We have for a long time been aware of a series of different adaptations in the structure of plants enabling them to endure excessive evaporation, and thus allowing them to live in place where the environment determines intense evaporation, or where the conditions of water absorption of the ground are unfavourable either physically or physiologically. Examples of such structures are: (1) covering of wax, (2) thick cuticle, (3) sub-epidermal protective tissue, (4) water tissue, (5) covering of hairs (6) covering of the stomata, (7) sinking of the stomata, (8) inclusion of the stomata in a space protected from air currents, (9) diminution of the evaporating surface, &c. The matter however is so complicated that it is very difficult to reach an exact appraisal of these adaptations in characterizing the individual plant communities biologically. ... In general we must content ourselves with showing the most frequently occurring adaptations, without going farther into the statistical investigation. ... A preliminary direct consideration of a series of evergreen phanerophytic communities, ... show that amongst the adaptations named, diminution of the transpiring surface, diminution in leaf size, is one of the adaptations generally in evidence; and since this adaptation is easy to observe and comparatively easy to measure, it is convenient to begin with it if we wish to use the statistical method on this domain.
Raunkiaer used the following size classes: *Leptophyll: less than 25 square millimetres *Nanophyll: 25–225 square millimetres *Microphyll: 225-2,025 square millimetres *Mesophyll: 2,025-18,225 square millimetres *Macrophyll: 18,225-164,025 square millimetres *Megaphyll: greater than 164,025 square millimetres Later authors have modified the classes and have sometimes used leaf length as a simpler measure than leaf area if the leaf shape is approximately an ellipse. For example, L.J. Webb Figure 2 used size classes: *Microphyll: less than 2,025 square millimetres *Notophyll: 2,025–4,500 square millimetres *Mesophyll: greater than 4,500 square millimetres


Examples of definitions


Single vegetable organisms with large leaves

*''
Gunnera manicata ''Gunnera manicata'', known as Brazilian giant-rhubarb or giant rhubarb, is a species of flowering plant in the family Gunneraceae from the coastal Serra do Mar Mountains of Santa Catarina, Parana and Rio Grande do Sul States, Brazil. In cultiva ...
'', giant ornamental rhubarb; leaves ; *''
Raphia regalis ''Raphia regalis'' is a species of flowering plant in the Palm Family Arecaceae r Palmae. It is found in Angola, Cameroon, Republic of the Congo, Gabon, and Nigeria. Its natural habitat is subtropical or tropical moist lowland forests. It is thre ...
'', composed leaves ; *''
Manicaria saccifera ''Manicaria'' (Common names Monkey Cap Palm or Bussu) is a Arecaceae, palm genus which is found in Trinidad, Central America, Central and South America. It contains two recognized species: #''Manicaria martiana'' Burret – Colombia, northweste ...
'', Amazonian palm; partially composed leaves ; *''
Marojejya darianii ''Marojejya darianii'', the big-leaf palm or Ravimbe Palm, is a species of flowering palm tree in the Palm Family (Arecaceae or Palmae). It is found only in the rainforests of Madagascar and was completely unknown to the larger world until 1984 ...
'', big-leaf palm; leaves ; *''
Johannesteijsmannia altifrons ''Johannesteijsmannia altifrons'' is a species of flowering plant in the palm family. It is native to southern Thailand, Peninsular Malaysia, Borneo, and Sumatra. Flowers of this species smell of sour milk. Description This is a medium-sized, ...
'', Joey palm; undivided leaves long; *''
Amorphophallus titanum The titan arum (''Amorphophallus titanum'') is a flowering plant in the family Araceae. It has a large unbranched inflorescence; a tall single leaf, branched like a tree; and a heavy tuber which enables the plant to produce the inflorescence. ' ...
'', titan arum; leaves area ; *''
Victoria amazonica ''Victoria amazonica'' is a species of flowering plant, the second largest in the water lily family Nymphaeaceae. It is called Vitória-Régia or Iaupê-Jaçanã ("the Jacanidae, jacana's waterlily") in Brazil and Atun Sisac ("great flower") in ...
'', giant Amazonian waterlily; aquatic plant with leaves long; leaves area .


See also

*
Leaf A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leav ...


References

{{reflist Ecological metrics Leaf morphology