''Nothotsuga'' is a genus of

coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

s in the family

Pinaceae The Pinaceae (), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, Pinyon_pine, piñons, larches, pines and spruces. The family is incl ...

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

. ''Nothotsuga'' contains only one living species, ''Nothotsuga longibracteata'', commonly known as the bristlecone hemlock or chang bao tie shan (), which is found in southeastern

, in southern

Fujian Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...

, northern

Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...

, northeast

Guangxi Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...

, northeast

Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...

, and southwest

Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...

. The genus was more diverse in the past, with its earliest fossils being known from Europe during the late

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

epoch, with the genus being present in Europe as recently as the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

epoch.

Description

''N. longibracteata'' is an

evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...

tree reaching tall. The leaves are flat, needle-like, long and broad, very similar to those of ''Tsuga''. The cones are very similar to those of ''Keteleeria'', but smaller, long, erect, and mature in about 6–8 months after

pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...

Taxonomy

In many respects, ''Nothotsuga'' is intermediate between the genera '' Keteleeria'' and ''

Tsuga ''Tsuga'' (, from Japanese (), the name of '' Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The English-language common name "hemlock" arose from a perceived similarity in the smell of it ...

''. It was discovered in 1932, and at first treated as ''Tsuga longibracteata'', being classified in its own

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

in 1989 when new research indicated how distinct it is from other species of ''Tsuga'' - by the larger, erect

cones In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines, ...

with exserted

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

s, and (like ''Keteleeria'') male cones in

umbel UMBEL (Upper Mapping and Binding Exchange Layer) is a logically organized knowledge graph of 34,000 concepts and entity types that can be used in information science for relating information from disparate sources to one another. It was retired ...

s, and from ''Keteleeria'' by the shorter

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

and smaller cones.

Conservation

It is a very rare tree listed as a

near-threatened species A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to endangerment in the near future, but it does not currently qu ...

by the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...

due to historical

deforestation Deforestation or forest clearance is the removal and destruction of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. Ab ...

, though it is now protected.

References

External links

Arboretum de Villardebelle - photo of cones

Gymnosperm Database

Flora of China
{{Taxonbar, from=Q63255706, from2=Q135182 Pinaceae Endemic flora of China Trees of China Flora of Fujian Flora of Guangdong Flora of Guangxi Flora of Guizhou Flora of Hunan Monotypic conifer genera Near threatened flora of Asia Plants described in 1932

Description

Taxonomy

Conservation

References

Further reading

External links