Northern Pygmy Owl
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The northern pygmy owl (''Glaucidium californicum'') is a small
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
native to western
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
.


Taxonomy

Some taxonomic authorities, including the
International Ornithologists' Union The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", ...
, separate this species from the
mountain pygmy owl The mountain pygmy owl or northern pygmy owl (''Glaucidium gnoma'') is a small species of owl from the family Strigidae. They reside throughout southern Arizona, New Mexico and Mexico. There is current taxonomic debate regarding its classificati ...
, the
Baja pygmy owl The Baja pygmy owl (''Glaucidium hoskinsii'') or cape pygmy owl, is a subspecies of northern pygmy owl restricted to the Mexican state of Baja California Sur. Although some taxonomists, including the International Ornithologists' Union, conside ...
, and the
Guatemalan pygmy owl The Guatemalan pygmy owl (''Glaucidium cobanense'') is a small "typical owl" in subfamily Surniinae. However, some taxonomic systems consider it to be a subspecies of northern pygmy owl (''G. gnoma'').HBW and BirdLife International (2020) ''Han ...
, while others, such as the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
, do not recognize the split and consider this bird conspecific with the group, with the northern pygmy owl taking the English name for them all. Furthermore, if the group is considered conspecific, ''G. gnoma'' becomes the scientific name because it is older. Clear differences in the territorial calls by males are the basis for the proposed split, with birds in the high elevations of Arizona and Mexico giving a two-note call while their more northerly congeners give a repeated single-note call. Results from
DNA sequence A nucleic acid sequence is a succession of bases within the nucleotides forming alleles within a DNA (using GACT) or RNA (GACU) molecule. This succession is denoted by a series of a set of five different letters that indicate the order of the nu ...
comparisons of cytochrome-b have been weak and inconclusive despite being referenced repeatedly as a justification for taxonomic splitting.


Subspecies

There are four recognized
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
: * ''G. c. californicum'' ( PL Sclater, 1857) – Pacific pygmy owl – central
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
to southwestern United States and northwestern Mexico * ''G. c. grinnelli'' ( Ridgway, 1914) – Coastal pygmy owl – coniferous forests of southeastern
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
to northern
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
* ''G. c. pinicola'' (
Nelson Nelson may refer to: Arts and entertainment * ''Nelson'' (1918 film), a historical film directed by Maurice Elvey * ''Nelson'' (1926 film), a historical film directed by Walter Summers * ''Nelson'' (opera), an opera by Lennox Berkeley to a lib ...
, 1910) – Rocky Mountain pygmy owl –
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
in west central United States * ''G. c. swarthi'' ( Grinnell, 1913) – Vancouver Island pygmy owl –
Vancouver Island Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian province of British Columbia. The island is in length, in width at its widest point, and in total area, while are of land. The island is the largest ...
in
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...


Description

Adults are in overall length and are gray, brownish-gray or
rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...
in colour. This owl has a round white-spotted head, weakly defined
facial disc In ornithology, the facial disc is the concave collection of feathers on the face of some birds—most notably owls—surrounding the eyes. The concavity of the facial disc forms a circular paraboloid that collects sound waves and directs those ...
, and dark upper breast, wings and tail, the latter quite long compared to other owls. The eyes are yellow and the bill is yellowish-green. The bird has two black nape spots outlined in white on the back of its head, which look like eyes. The mid to lower breast is white with darker vertical streaking. Legs are feathered down to the four well-armed toes on each foot.


Distribution and habitat

The northern pygmy owl is native to
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, and
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. Their habitat includes temperate, subtropical and tropical moist forest, savanna, and wetlands. In Oregon and Washington they are known to nest and forage in the center of dense, continuous forests, near streams. An example of their habitat is
Forest Park A forest park is a park whose main theme is its forest of trees. Forest parks are found both in the mountains and in the urban environment. Examples Chile * Forest Park, Santiago China * Gongqing Forest Park, Shanghai * Mufushan National Fore ...
in
Portland, Oregon Portland ( ) is the List of cities in Oregon, most populous city in the U.S. state of Oregon, located in the Pacific Northwest region. Situated close to northwest Oregon at the confluence of the Willamette River, Willamette and Columbia River, ...
, USA. Their breeding habitat includes open to semi-open woodlands of foothills and mountains in western
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
.


Behavior

Males will regularly perch at the top of the tallest available conifer trees to issue their territorial call, making them somewhat ventriloquistic in sloped landscapes, and causing distress and confusion among observers on the ground hoping to get a glimpse. They are incredibly hard to spot because of their size and color.


Breeding

They usually nest in a tree cavity and will often use old
woodpecker Woodpeckers are part of the bird family (biology), family Picidae, which also includes the piculets, wrynecks and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar and the extreme ...
holes. The female lays 2–7 eggs, typically 4–6. Nest tree species may include
Douglas fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Or ...
,
western redcedar ''Thuja plicata'' is a large evergreen coniferous tree in the family Cupressaceae, native to the Pacific Northwest of North America. Its common name is western redcedar in the U.S. or western red cedar in the UK, and it is also called pacific re ...
,
western hemlock ''Tsuga heterophylla'', the western hemlock or western hemlock-spruce, is a species of hemlock native to the northwest coast of North America, with its northwestern limit on the Kenai Peninsula, Alaska, and its southeastern limit in northern ...
and
red alder ''Alnus rubra'', the red alder, is a deciduous broadleaf tree native to western North America (Alaska, Yukon, British Columbia, Washington, Oregon, California, Idaho and Montana). Description ''Alnus rubra'' is the largest species of alder in ...
. Early in the breeding cycle males establish and defend a territory of perhaps 250 hectares (about 1 sq. mi.).Giese, A (2003). "Breeding Season Habitat Use and Ecology of Male Mountain Pygmy Owls". ''Journal of Raptor Research''. During the breeding cycle the female incubates the eggs, broods the young and guards the nest. The male hunts, making food deliveries approximately every 2 hours. The male must feed his mate, the young (typically 5) and himself. The male hunts from dawn to dusk as the young near fledging, and during the first weeks after they leave the nest. The young leave the nest (fledge) by making an initial flight that may be a short hop to a nearby branch, or an explosive burst into an adjacent tree where they land by grasping whatever branch is first contacted, sometimes clinging upside-down. Owls at this stage are sometimes called "branchers" for their clinging, dangling and climbing behaviors. The second day after fledging, the young gradually climb and fly upward into the forest canopy, where they spend their first few weeks, at times perched "shoulder-to-shoulder" with their siblings, begging for food. Despite many statements in popular literature, no reliable information exists on the seasonal movements of this species. It is not known whether these pygmy-owls maintain the same territory or same mate year to year, though these questions are being investigated. Dispersal of young and influences on their mortality are also poorly known, though
barred owl The barred owl (''Strix varia''), also known as the northern barred owl, striped owl or, more informally, hoot owl or eight-hooter owl, is a North American large species of owl. A member of the true owl family, Strigidae, they belong to the genus ...
s and
spotted owl The spotted owl (''Strix occidentalis'') is a species of true owl. It is a resident species of old-growth forests in western North America, where it nests in tree hollows, old bird of prey nests, or rock crevices. Nests can be between high a ...
s are known to prey on pygmy owls.


Feeding

Pygmy owls are purportedly "sit-and-wait" predators, though they in fact hunt somewhat actively, moving from perch to perch with short flights, and pursuing prey at all levels of forest structure. They swoop down on prey; they may also catch insects in flight. They eat small mammals, birds and large insects, and may take a variety of other vertebrates and invertebrates. Mountain pygmy owls occasionally take prey species the same size or larger than themselves (e.g.
California quail The California quail (''Callipepla californica''), also known as the California valley quail or Valley quail, is a small ground-dwelling bird in the New World quail family. These birds have a curving crest, '' plume'' or topknot made of six fea ...
); however, small to medium-sized birds and small mammals are the norm. They've been observed eating
Wilson's warbler Wilson's warbler (''Cardellina pusilla'') is a small New World warbler. It is greenish above and yellow below, with rounded wings and a long, slim tail. The male has a black crown patch; depending on the subspecies, that mark is reduced or absent ...
s. These owls are diurnal, and also active at dawn and dusk.


References

* ''Field Guide to the Birds of North America''. National Geographic. . * Josep del Hoyo, ed. ''Handbook of the Birds of the World Vol 5''. . * David Allen Sibley. ''The Sibley Guide to Birds''. National Audubon Society. .


External links


(Northern) Pygmy Owls
Documentary produced by
Oregon Field Guide ''Oregon Field Guide'' is a weekly television program produced by Oregon Public Broadcasting focusing on recreation, the outdoors, and environmental issues in the state of Oregon. The show has become part of the Oregon zeitgeist. Steve Amen is th ...
. {{Taxonbar, from=Q2963132
northern pygmy owl The northern pygmy owl (''Glaucidium californicum'') is a small owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad ...
Native birds of Western Canada Native birds of the Western United States
northern pygmy owl The northern pygmy owl (''Glaucidium californicum'') is a small owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad ...
northern pygmy owl The northern pygmy owl (''Glaucidium californicum'') is a small owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad ...
northern pygmy owl The northern pygmy owl (''Glaucidium californicum'') is a small owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad ...