The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a
bird of prey
Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
. It breeds throughout the northern parts of the
northern hemisphere
The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
in
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
and the northernmost
USA
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 states and a federal capital district, Washington, D.C. The 48 contiguous ...
.
The northern harrier
migrates south in winter, with breeding birds in Canada and northern Great Plains of the U.S. moving to the American south, Mexico, and
Central America
Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
. In the midwestern, mountain west, and north Atlantic states of the U.S., they may be present all year. This bird inhabits prairies, open areas, and marshes.
Taxonomy
In 1750 the English naturalist
George Edwards included an illustration and a description of the northern harrier in the third volume of his ''A Natural History of Uncommon Birds''. He used the English name "The Ring-tail'd Hawk". Edwards based his hand-coloured etching on a bird collected near the
Hudson Bay
Hudson Bay, sometimes called Hudson's Bay (usually historically), is a large body of Saline water, saltwater in northeastern Canada with a surface area of . It is located north of Ontario, west of Quebec, northeast of Manitoba, and southeast o ...
in Canada and brought to London by
James Isham James Isham (1716–1761) was chief factor (master) at both York Factory and Prince of Wales Fort, Fort Prince of Wales in Canada during the mid-1700s. He kept detailed journals that described life in the region, including flora and fauna that were ...
. When in 1766 the Swedish naturalist
Carl Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
updated his ''
Systema Naturae
' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' for the
twelfth edition, he placed the northern harrier with the falcons and eagles in the
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Falco''. Linnaeus included a brief description, coined the
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Falco hudsonius'' and cited Edwards' work. The northern harrier is now placed in the genus ''
Circus
A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicy ...
'', introduced by the French naturalist
Bernard Germain de Lacépède
Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French natural history, naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's g ...
in 1799.
The genus name ''Circus'' is derived from the
Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''kirkos'', referring to a bird of prey named for its circling flight (''kirkos'', "circle"). The specific epithet ''hudsonius'' is from "Hudson Bay", the
type locality. The species is
monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.
[
The northern harrier was formerly considered to be ]conspecific
Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species.
Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
with the hen harrier
The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Palearctic, Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl.
It bird migration, migrates to more southerly areas in winter. Eurasian ...
.[
]
Description
The northern harrier is [ long with a wingspan. It resembles other harriers in having distinct male and female plumages. The sexes also differ in weight, with males weighing , with an average of , and females weighing , with an average of .][ Among standard measurements, the wing chord is , the ]tail
The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
is and the tarsus is .[ It is relatively long-winged and long-tailed, having the longest wing and tail relative to its body size of any raptor occurring in North America.][
The northern harrier breeds in North America, and its closest relative is the cinereous harrier (''C. cinereus'') of South America. The male's plumage is darker grey than that of the hen harrier, and the female is also darker and more rufous.][ The adult male is sometimes nicknamed the "Grey Ghost", because of his striking plumage and spectral aura.]
The northern harrier is mostly silent, although males and females will both give fast ''kek'' notes that will last 1-2 seconds. When faced with predators or surrounded by smaller birds, they still emit ''kek'' notes but at a higher pitch. During the mating season, the female emits a loud scream, which causes the male to either mate or provide food.
Behaviour
This medium-sized raptor
Raptor(s) or RAPTOR may refer to:
Animals
The word "raptor" refers to several groups of avian and non-avian dinosaurs which primarily capture and subdue/kill prey with their talons.
* Raptor (bird) or bird of prey, a bird that primarily hunt ...
breeds on moorland
Moorland or moor is a type of Habitat (ecology), habitat found in upland (geology), upland areas in temperate grasslands, savannas, and shrublands and the biomes of montane grasslands and shrublands, characterised by low-growing vegetation on So ...
, bog
A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and musk ...
s, prairie
Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
s, farm
A farm (also called an agricultural holding) is an area of land that is devoted primarily to agricultural processes with the primary objective of producing food and other crops; it is the basic facility in food production. The name is used fo ...
land coastal prairies, marsh
In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...
es, grassland
A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s, swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and other assorted open areas.[ A male will maintain a territory averaging , though male territories have ranged from .][
]
These are one of the few raptorial birds known to practice polygyny
Polygyny () is a form of polygamy entailing the marriage of a man to several women. The term polygyny is from Neoclassical Greek πολυγυνία (); .
Incidence
Polygyny is more widespread in Africa than in any other continent. Some scholar ...
– one male mates with several females. Up to five females have been known to mate with one male in a season.[ The nest is built on the ground or on a mound of dirt or vegetation. Nests are made of sticks and are lined inside with grass and leaves. Four to eight (exceptionally 2 to 10) whitish ]eggs
An egg is an organic vessel in which an embryo begins to develop.
Egg, EGG or eggs may also refer to:
Biology
* Egg cell, the female reproductive cell (gamete) in oogamous organisms
Food
* Eggs as food
Places
* Egg, Austria
* Egg, Switzerland ...
are laid.[ The eggs measure approximately .][ The eggs are incubated mostly by the female for 31 to 32 days. When incubating eggs, the female sits on the nest while the male hunts and brings food to her and the chicks.][ The male will help feed chicks after they hatch, but does not usually watch them for a greater period of time than around 5 minutes.][ The male usually passes off food to the female, which she then feeds to the young, although later the female will capture food and simply drop into the nest for her nestlings to eat.][ The chicks fledge at around 36 days old, though breeding maturity is not reached until 2 years in females and 3 years in males.
]
Hunting behavior
This is a typical harrier, which hunts on long wings held in a shallow V in its low flight during which the bird closely hugs the contours of the land below it. Northern or hen harriers hunt primarily small mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s, as do most harriers. Preferred prey species can include vole
Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s, cotton rats and ground squirrel
Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s. Up to 95% of the diet comprises small mammals.[ However, ]bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s are hunted with some regularity as well, especially by males. Preferred avian prey include passerine
A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
s of open country (i.e. sparrows, lark
Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
s, pipit
The pipits are a cosmopolitan genus, ''Anthus'', of small passerine birds with medium to long tails. Along with the wagtails and longclaws, the pipits make up the family Motacillidae. The genus is widespread, occurring across most of the world, ...
s), small shorebird
245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots
Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s and the young of waterfowl
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...
and galliforms
Galliformes is an order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed disperser ...
. Supplementing the diet occasionally are amphibian
Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s (especially frog
A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s), reptile
Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s and insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s (especially orthoptera
Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – gras ...
ns).[ The species has been observed to hunt ]bat
Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s if these are available. Larger prey such as rabbit
Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s and adult duck
Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
s are taken sometimes and harriers have been known to subdue these by drowning them in water.[ Harriers hunt by surprising prey while flying low to the ground in open areas, as they drift low over fields and moors.][ The harriers circle an area several times listening and looking for prey.][ Harriers use hearing regularly to find prey, as they have exceptionally good hearing for diurnal raptors, this being the function of their ]owl
Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
-like facial disc.[ This harrier tends to be a very vocal bird while it glides over its hunting ground.
]
Mortality and competition
Little information is available on longevity in northern harriers. The longest lived known bird is 16 years and 5 months. However, adults rarely live more than 8 years. Early mortality mainly results from predation. Predators of eggs and nestlings include raccoon
The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...
s, skunk
Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
s, badger
Badgers are medium-sized short-legged omnivores in the superfamily Musteloidea. Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by their squat bodies and adaptions for fossorial activity rather than by the ...
s, fox
Foxes are small-to-medium-sized omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull; upright, triangular ears; a pointed, slightly upturned snout; and a long, bushy tail ("brush").
Twelve species ...
es, crows and ravens, dog
The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s, and owl
Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s. Fledglings are also predated regularly, especially by great horned owl
The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air") or the hoot owl, is a large owl native to the Americas. It is an extreme ...
s.[ Both parents attack potential predators with alarm calls and striking with talons. ]Short-eared owl
The short-eared owl (''Asio flammeus'') is a widespread grassland species in the family Strigidae. Owls belonging to genus ''Asio'' are known as the eared owls, as they have tufts of feathers resembling mammalian ears. These "ear" tufts may or ...
s are natural competitors of this species that favor the same prey and habitat, as well as having a similarly broad distribution. Occasionally, both harriers and short-eared owls will harass each other until the victim drops its prey and it can be stolen, a practice known as kleptoparasitism
Kleptoparasitism (originally spelt clepto-parasitism, meaning "parasitism by theft") is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct f ...
. Most commonly, the harriers are the aggressors pirating prey from owls.[
]
Status
This species has a large range, and there is evidence of a population decline, but the species is not believed to approach the thresholds for the population decline criterion of the IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
(i.e., declining more than 30% in ten years or three generations). It is therefore classified as "least concern".
Relationship with humans
Some Native American tribes believe that seeing a hawk on your wedding day is a sign of a long, happy marriage. Unlike many raptors, hen or northern harriers have historically been favorably regarded by farmers because they eat mice that damage crops and predators of quail eggs. Harriers are sometimes called "good hawks" because they pose no threat to poultry as some hawks do. Heavy pesticide use in the 1970s and 1980s caused a decline in harrier populations.[
]
References
External links
Northern Harrier Species Account
– Cornell Lab of Ornithology
*
{{Taxonbar, from=Q2603050
northern harrier
The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost United States, USA.
The northern ...
Birds of prey of North America
Birds of the Dominican Republic
northern harrier
The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost United States, USA.
The northern ...
northern harrier
The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost United States, USA.
The northern ...