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''Nopcsaspondylus'' (meaning " Nopcsa's
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
", in reference to the original describer) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
rebbachisaurid Rebbachisauridae is a Family (biology), family of sauropod dinosaurs known from fragmentary fossil remains from the Cretaceous of South America, Africa, North America, Europe and possibly Central Asia. Taxonomy In 1990 sauropod specialist Jack M ...
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
(a type of large, long-necked
quadruped Quadrupedalism is a form of locomotion in which animals have four legs that are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four legs is said to be a quadruped (fr ...
al
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
dinosaur) from the
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
-age (
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
)
Candeleros Formation The Candeleros Formation is a geologic formation that outcrop, crops out in the Rio Negro Province, Río Negro, Neuquen Province, Neuquén, and Mendoza Province, Mendoza provinces of northern Patagonia, Argentina. It is the oldest formation in the ...
of
Neuquén Neuquén (; ) is the capital city of the Argentine province of Neuquén and of the Confluencia Department, located in the east of the province. It occupies a strip of land west of the confluence of the Limay and Neuquén rivers which form t ...
,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. It is based on a now-lost back vertebra described by Nopcsa in 1902 but not named at the time. The specimen had a small
vertebral body Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
and large hollows, now known to be typical of rebbachisaurids.


History of research

The only known specimen of ''Nopcsaspondylus'' is a
fossilized A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
dorsal vertebra In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebrae of intermediate size between the cervical and lumbar ve ...
collected in 1889 by Slovenian researcher
Hugo Zapałowicz Hugo Zapałowicz (15 November 1852, in Ljubljana – 20 November 1917, in Perovsk Kyzylorda.html" ;"title="ow Kyzylorda">ow Kyzylorda/nowiki>, Kazakhstan) was a botanist, natural scientist, traveller, and military lawyer. Zapałowicz was a pion ...
and transferred to the geological collection of the
University of Vienna The University of Vienna (, ) is a public university, public research university in Vienna, Austria. Founded by Rudolf IV, Duke of Austria, Duke Rudolph IV in 1365, it is the oldest university in the German-speaking world and among the largest ...
. The first person known to study the specimen, Hungarian paleontologist
Franz Nopcsa von Felső-Szilvás Baron Franz Nopcsa von Felső-Szilvás (also Baron Nopcsa von Felső-Szilvás, Baron Nopcsa, Ferenc Nopcsa, báró felsőszilvási Nopcsa Ferenc, Baron Franz Nopcsa, and Franz Baron Nopcsa; May 3, 1877 – April 25, 1933) was a Hungarian Aristoc ...
, described it in a 1902 publication and recognized it as belonging to a
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur, assigning it to the genus ''
Bothriospondylus ''Bothriospondylus'', from Ancient Greek βοθρίον (''bothríon''), meaning "trench", and σπόνδυλος (''spóndulos''), meaning "vertebra", is a nomen dubium, dubious genus of neosauropod sauropod dinosaur. It lived during the Late Ju ...
''. According to Nopcsa, the label on the fossil stated that it was collected on the left bank of the
Limay River The Limay River is an important river in the northwestern Argentine Patagonia (the region of Comahue). It originates at the eastern end of the Nahuel Huapi Lake and flows in a meandering path for about , collecting the waters of several tributa ...
in
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
, about 80 kilometers south of the
confluence In geography, a confluence (also ''conflux'') occurs where two or more watercourses join to form a single channel (geography), channel. A confluence can occur in several configurations: at the point where a tributary joins a larger river (main ...
of the Limay and
Neuquén Neuquén (; ) is the capital city of the Argentine province of Neuquén and of the Confluencia Department, located in the east of the province. It occupies a strip of land west of the confluence of the Limay and Neuquén rivers which form t ...
rivers. A year later, American paleontologist
John Bell Hatcher John Bell Hatcher (October 11, 1861 – July 3, 1904) was an American paleontologist and fossil hunter known as the "king of collectors" and best known for discovering ''Torosaurus'' and ''Triceratops'', two genera of dinosaurs described by O ...
claimed that this vertebra was indistinguishable from those of ''
Haplocanthosaurus ''Haplocanthosaurus'' (meaning "simple spined lizard") is a genus of diplodocoid sauropod dinosaur. Two species, ''H. delfsi'' and ''H. priscus'', are known from incomplete fossil skeletons. They lived during the Late Jurassic (Kimmeridgian sta ...
'', a sauropod known from
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
, and that while the Argentine vertebra does not undoubtedly represent the same genus as the Colorado material, the two are very similar in structure. The vertebra would next be studied by German paleontologist
Friedrich von Huene Baron Friedrich Richard von Hoyningen-Huene (22 March 1875 – 4 April 1969) was a German nobleman paleontologist who described a large number of dinosaurs, more than anyone else in 20th-century Europe. He studied a range of Permo-Carbonife ...
, who wrote a
monograph A monograph is generally a long-form work on one (usually scholarly) subject, or one aspect of a subject, typically created by a single author or artist (or, sometimes, by two or more authors). Traditionally it is in written form and published a ...
published in 1929 in which he reassigns the material to the species '' Titanosaurus australis'', and the locality from which it originates is named as Alarcón for the first time. The ''Nopcsaspondylus'' specimen would not be published on again until the release of ''
The Dinosauria ''The Dinosauria'' is an encyclopedia on dinosaurs, edited by paleontologists David B. Weishampel, Peter Dodson, and Halszka Osmólska. It has been published in two editions by the University of California Press, with the first edition in 1990 ...
'' in 1990, in which the vertebra was deemed similar to those found in northern Africa of '' Rebbachisaurus garasbae'', supposedly extending the range of the ''Rebbachisaurus'' genus into South America as well. Five years afterwards, Argentine paleontologists
Jorge O. Calvo Jorge Orlando Calvo (27 April 1961 – 10 January 2023)Leonardo Salgado Leonardo Salgado is an Argentine palaeontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossi ...
described a new species which they named '' Rebbachisaurus tessonei'' (currently moved to the genus ''
Limaysaurus ''Limaysaurus'' ("Limay lizard") is a genus represented by a single species of rebbachisaurid sauropod dinosaurs, which lived during the mid-Cretaceous period, about 99.6 to 94 million years ago, in the Cenomanian, in what is now South America (n ...
''), the specimens of which originate from the same locality (now named as Barda Alarcón) as the vertebra first described by Nopcsa 93 years prior. They therefore claimed that Nopcsa's specimen, at this point already considered lost, was a vertebra of ''R. tessonei''. In 2007, Argentine paleontologist Sebastián Apesteguía became the first author to recognize that the vertebra described by Nopcsa represents a new genus and species, and thus named this species as ''Nopcsaspondylus alarconensis''. The generic name combines the surname of the first researcher to study it with the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
word (, meaning "vertebra"), while the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
refers to Barda Alarcón, the locality where the fossil was discovered.


References

{{Taxonbar, from=Q2102851 Rebbachisauridae Dinosaur genera Cenomanian dinosaurs Candeleros Formation Fossil taxa described in 2007 Dinosaurs of Argentina Taxa with lost type specimens