Nodosaurus
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''Nodosaurus'' (meaning 'knobbed lizard') is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
nodosaurid Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the s ...
ankylosaurian Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
, the fossils of which are found exclusively in the Frontier Formation in
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
.


Description

''Nodosaurus'' grew up to roughly long and it was an
ornithischian Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
dinosaur with bony dermal plates covering the top of its body, and it may have had spikes along its side as well. The dermal plates were arranged in bands along its body, with narrow bands over the ribs alternating with wider plates in between. These wider plates were covered in regularly arranged bony nodules, which give the animal its scientific name. In 2010
Paul Paul may refer to: People * Paul (given name), a given name, including a list of people * Paul (surname), a list of people * Paul the Apostle, an apostle who wrote many of the books of the New Testament * Ray Hildebrand, half of the singing duo ...
estimated its length at 6 meters (20 ft) and its weight at 3.5 tonnes (3.85 short tons). It had four short legs, five-toed feet, a short neck, and a long, stiff, clubless tail. The head was narrow, with a pointed snout, powerful jaws, and small teeth. It perhaps ate soft plants, as it would have been unable to chew tough, fibrous ones; or alternatively it may have processed the latter with
gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In ...
s and its enormous intestinal apparatus.


History and classification

During the
Bone Wars The Bone Wars, also known as the Great Dinosaur Rush, was a period of intense and ruthlessly competitive fossil hunting and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope (of the Aca ...
between palaeontologists
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
and
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontology, paleontologist, comparative anatomy, comparative anatomist, herpetology, herpetologist, and ichthyology, ichthyologist. Born to a wealthy Quaker fam ...
, Marsh sent his collector
William Harlow Reed William Harlow Reed (9 June 1848 – 24 April 1915) was an American fossil collector and pioneer. He served as a curator at the Museum of Geology at the University of Wyoming, Laramie. He collected for a while for Othniel Charles Marsh but left a ...
to the
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
strata of the Frontier Formation of Albany County, Wyoming to collect fossil mammals and reptiles. Reed collected a partial postcranial skeleton (YPM VP 1815) on July 17, 1881 southeast of the productive Quarry 13, the skeleton including: 3 dorsal and 13 caudal vertebrae, 3 dorsal ribs, fragmentary forelimbs, a partial pelvis, femora, tibiae, partial left pes, and several osteoderms.Raven, T. J. (2021). ''The Taxonomic, Phylogenetic, Biogeographic and Macroevolutionary History of the Armoured Dinosaurs (Ornithischia: Thyreophora)''(Doctoral dissertation, University of Brighton).Coombs, Jr. (1978). "The families of the ornithischian dinosaur order Ankylosauria". Palaeontology. 21 (1): 143–170. One of the first armored dinosaurs to be discovered in North America, ''Nodosaurus textilis'' was named by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
in 1889, the generic name meaning "knobbed lizard" and the specific name meaning "woven".Lull, R. S. (1921)
The Cretaceous armored dinosaur, Nodosaurus textilis Marsh.
/ref> Marsh assigned the genus to
Stegosauria Stegosauria is a group of Herbivore, herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous Period (geology), periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere (North America, Europe a ...
, but later assigned it to its own family,
Nodosauridae Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the ...
, in 1890 based on the heavy dermal armor, solid bones, large forelimbs, and ungulate feet. The type specimen remains the only definite specimen of ''Nodosaurus'', though ''
Stegopelta ''Stegopelta'' (meaning "roofed shield") is a genus of struthiosaurin nodosaurid dinosaur. It is based on a partial skeleton from the latest Albian- earliest Cenomanian-age Lower and Upper Cretaceous Belle Fourche Member of the Frontier Forma ...
'' has been considered a synonym in the past, it is most likely a distinct struthiosaurin. Richard Swan Lull did a more comprehensive description of ''Nodosaurus'' in 1921, who assigned the taxa ''
Stegopelta ''Stegopelta'' (meaning "roofed shield") is a genus of struthiosaurin nodosaurid dinosaur. It is based on a partial skeleton from the latest Albian- earliest Cenomanian-age Lower and Upper Cretaceous Belle Fourche Member of the Frontier Forma ...
,
Hoplitosaurus ''Hoplitosaurus'' (meaning "Hoplite lizard") was a genus of armored dinosaur related to ''Polacanthus''. It was named from a partial skeleton found in the Barremian-age Lower Cretaceous Lakota Formation of Custer County, South Dakota. It is an ...
,
Hierosaurus ''Hierosaurus'' (meaning "sacred lizard") is an extinct genus of Nodosauridae, nodosaurid ankylosaur which lived during the Late Cretaceous 87 to 82 million years ago. Its fossils were found in the Smoky Hill Chalk, Smoky Hill Chalk Member of the ...
,
Ankylosaurus ''Ankylosaurus'' is a genus of Thyreophora, armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period (geology), Period, about 68–66 million years ago, in western North America, m ...
,'' and placed the British ''
Polacanthus ''Polacanthus'', deriving its name from the Ancient Greek polys-/πολύς- "many" and akantha/ἄκανθα "thorn" or "prickle", is an early armoured, spiked, plant-eating ankylosaurian dinosaur from the early Cretaceous period of England. In ...
'' as a relative of the family''.'' Within Nodosauridae, ''Nodosaurus'' falls out in the subfamily Nodosaurinae, formally defined as the largest clade containing ''Nodosaurus textilis'' but not ''
Hylaeosaurus armatus ''Hylaeosaurus'' ( ; Greek: / "belonging to the forest" and / "lizard") is a herbivorous ankylosaurian dinosaur that lived about 136 million years ago, in the late Valanginian stage of the early Cretaceous period of England. It was found in ...
'', '' Mymoorapelta maysi'' and ''Polacanthus foxii''. The
2018 Events January * January 1 – Bulgaria takes over the Presidency of the Council of the European Union, after the Estonian presidency. * January 4 – SPLM-IO rebels loyal to Chan Garang Lual start a raid against Juba, capital of ...
phylogenetic analysis of Nodosauridae by Rivera-Sylva and colleagues is below.


See also

*
Timeline of ankylosaur research This timeline of ankylosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spik ...


References

{{Taxonbar, from=Q134209 Nodosauridae Dinosaur genera Cenomanian dinosaurs Taxa named by Othniel Charles Marsh Fossil taxa described in 1889 Dinosaurs of the United States