''Niassodon'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of kingoriid

dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, t ...

therapsid known from the

Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...

Niassa Province Niassa is a province of Mozambique with an area of 129,056 km2 and a population of 1,810,794 (2017). It is the most sparsely populated province in the country. Lichinga is the capital of the province. There are a minimum estimated 450,00 ...

, northern

Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...

. It contains a single species, ''Niassodon mfumukasi''.

Discovery and naming

''Niassodon'' was first described and named by Rui Castanhinha, Ricardo Araújo, Luís C. Júnior, Kenneth D. Angielczyk, Gabriel G. Martins, Rui M. S. Martins, Claudine Chaouiya, Felix Beckmann and Fabian Wilde in

2013 2013 was the first year since 1987 to contain four unique digits (a span of 26 years). 2013 was designated as: *International Year of Water Cooperation *International Year of Quinoa Events January * January 5 – 2013 Craig, Alask ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Niassodon mfumukasi''. The generic name combines the word ''Niassa'', meaning "

lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...

" in Chiyao and the name of the northwestern province in Mozambique where ''Niassodon'' was found, with ''odontos'', meaning "tooth" in

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''Mfumukasi'' means "queen" in

Nyanja Chewa ( ; also known as Nyanja ) is a Bantu language spoken in Malawi and a recognised minority in Zambia and Mozambique. The noun class prefix ''chi-'' is used for languages, so the language is often called or Chinyanja. In Malawi, the name wa ...

, in honor of the members of the local Nyanja

matriarchal Matriarchy is a social system in which positions of power and privilege are held by women. In a broader sense it can also extend to moral authority, social privilege, and control of property. While those definitions apply in general English, ...

society and all Mozambican women. ''Niassodon'' is known solely from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

ML1620, a partial

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

which has returned to the Museu Nacional de Geologia,

Maputo Maputo () is the Capital city, capital and largest city of Mozambique. Located near the southern end of the country, it is within of the borders with Eswatini and South Africa. The city has a population of 1,088,449 (as of 2017) distributed ov ...

, Mozambique. The holotype consists of a complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, a

mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

, series of 19

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...

, sacral and

caudal vertebrae Caudal vertebrae are the vertebrae of the tail in many vertebrates. In birds, the last few caudal vertebrae fuse into the pygostyle, and in apes, including humans, the caudal vertebrae are fused into the coccyx. In many reptiles, some of the caud ...

rib In vertebrate anatomy, ribs () are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the thoracic cavity, enabling the lungs to expand and thus facilitate breathing by expanding the ...

s, both ilia and a partial

femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...

, from a single small individual (skull length is 58 mm). It was discovered by Ricardo Araújo and collected during a 2009 expedition in the Metangula Graben, under the supervision of Projecto PaleoMoz. ML1620 came from the K5 Formation located near Tulo, a small village situated along the Metangula-Cóbue road. The fossil bed is composed of a grey mudstone with abundant septaria-like calcareous

concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...

s, dating to the

References

Dicynodontia Lopingian synapsids of Africa Lopingian genus first appearances Lopingian genus extinctions Fossils of Mozambique Fossil taxa described in 2013 Articles containing video clips Anomodont genera {{anomodont-stub