''Nesodactylus'' was a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

rhamphorhynchid Rhamphorhynchidae is a group of early pterosaurs named after ''Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study o ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...

from the middle-late Oxfordian ageDe la Fuente, M. S., & Iturralde-Vinent, M. (2001). A new pleurodiran turtle from the Jagua Formation (Oxfordian) of western Cuba. Journal Information, 75(4).

Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

Jagua Formation The Jagua Formation is a Late Jurassic (middle to late Oxfordian) geologic formation in the Sierra de los Órganos and Sierra del Rosario mountain ranges in Pinar del Río Province, western Cuba. Plesiosaur, pliosaur, pterosaur,Barrett et al., ...

Pinar del Río Pinar del Río is the capital city of Pinar del Río Province, Cuba. With a population of 139,336 (2004) in a municipality of 190,332, it is the List of cities in Cuba, 10th-largest city in Cuba. Inhabitants of the area are called ''Pinareños'' ...

, western

Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...

. Its remains were collected but not prepared by

Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...

in 1918, from rocks better known for their fossils of marine life. When seven black chalkstone blocks were prepared from 1966 by

Richard Lund Richard Lund (9 July 1885 – 27 September 1960) was a Swedish film and theatre actor. He made his stage debut at Stora Teatern in Gothenburg in 1904 and later appeared in more than 70 films between 1912 and 1952, making his most important ...

by dissolving the substrate in

acid In computer science, ACID ( atomicity, consistency, isolation, durability) is a set of properties of database transactions intended to guarantee data validity despite errors, power failures, and other mishaps. In the context of databases, a sequ ...

, this revealed the remains of a pterosaur. Ned Colbert described and named the genus in 1969. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Nesodactylus hesperius''. The genus name is derived from Greek ''nesos'', "island" and ''daktylos'', "finger", a reference to the island of Cuba and the typical wing finger of pterosaurs. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

means "western", from Greek ''hesperios''. The genus is based on

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

AMNH 2000, a partial skeleton including a skull fragment, numerous

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

e from all parts of the spine and

tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...

, zygapophyses (interpreted by Colbert as ossified

tendons A tendon or sinew is a tough, high-tensile-strength band of dense fibrous connective tissue that connects muscle to bone. It is able to transmit the mechanical forces of muscle contraction to the skeletal system without sacrificing its ability ...

) on the tail, the

pectoral girdle The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists of t ...

and a very deeply keeled

sternum The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sh ...

, arms and partial hands, part of the

pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

, parts of both

femora The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with t ...

, partial

metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...

s, and ribs. The specimen was disarticulated but associated and not very compressed; during the preparation from the

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

with acid, the bones were not completely removed. Colbert found ''Nesodactylus'' to have had longer wings and more robust limbs and longer legs than related ''

Rhamphorhynchus ''Rhamphorhynchus'' (, from Ancient Greek ''rhamphos'' meaning "beak" and ''rhynchus'' meaning "snout") is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such ...

'', although of a similar size and overall anatomy. He classified it as a

and more precisely as a member of the

Rhamphorhynchinae Rhamphorhynchidae is a group of early pterosaurs named after ''Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study o ...

. In 1977

James A. Jensen James Alvin Jensen (August 2, 1918 – December 14, 1998), was an American paleontologist. His extensive collecting program at Brigham Young University in the Utah-Colorado region which spanned 23 years was comparable in terms of the number of ...

and

John Ostrom John Harold Ostrom (February 18, 1928 – July 16, 2005) was an American paleontologist who revolutionized modern understanding of dinosaurs in the 1960s. As first proposed by Thomas Henry Huxley in the 1860s, Ostrom showed that dinosaurs were ...

by mistake referred to it as ''Nesodon'' (1977). Although there is little overlapping material with contemporaneous '' Cacibupteryx'', the two are clearly different based on details of the

elbow The elbow is the region between the arm and the forearm that surrounds the elbow joint. The elbow includes prominent landmarks such as the olecranon, the cubital fossa (also called the chelidon, or the elbow pit), and the lateral and the media ...

and

quadrate Quadrate may refer to: * Quadrate bone * Quadrate (heraldry) * Quadrate lobe of liver * Quadrate tubercle The quadrate tubercle is a small tubercle found upon the upper part of the femur. It serves as a point of insertion of the quadratus femori ...

. At least one recent review suggests it was a rhamphorhynchine, while another does not classify it.

References

{{Taxonbar, from=Q383964 Rhamphorhynchids Late Jurassic pterosaurs of North America Oxfordian life Jurassic Cuba Fossils of Cuba Fossil taxa described in 1969 Taxa named by Edwin H. Colbert

See also

References