''Neottia ovata'' (formerly ''Listera ovata''), the common twayblade or eggleaf twayblade, is a terrestrial orchid widespread across much of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and

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Description

The flowering stems are typically tall, occasionally up to . There are two large opposite basal leaves, long. A variable number of flowers is borne on the stems, usually more than 15 but less than 100. The flowers are small and yellowish-green in colour. The

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s and the two side

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s form a fairly open hood, 5–6 mm (0.2 in) long; the labellum or lip (the central petal) is long and is divided at the end into two lobes. Due to its slender profile, small flowers and green colour this species can be hard to spot.

Distribution and habitat

Found across much of Europe including the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

, as well as

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...

, Southwest Asia and the

Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...

. It has been introduced into

Ontario Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...

Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...

, where it has been called the eggleaf twayblade. (as ''Listera ovata'') ''Neottia ovata'' grows in a variety of habitats: woods, meadows, dune-slacks and moorland.

Ecology

The flower of ''Neottia ovata'' is well accessible for a wide range of insects. It is above all pollinated by parasitic

wasps A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...

sawflies Sawflies are wasp-like insects that are in the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plant ...

and

beetles Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

. The pollinia lie free on top of the gutter-shaped rostellum, an organ that is filled with viscid fluid. When an insect touches the sensitive tip of the rostellum, the viscid fluid is ejected and glues the pollinia to the visitor's body. Fruit set is quite high. Over 60 species of mycorrhizal fungi have been indicated to form associations with ''N. ovata''. ''Neottia ovata'' is one of the most common European orchid species, and this is partially explained by it being less species-selective both in terms of pollinators and mycorrhizal partners. It is also an inconspicuous species, blending in to vegetation, which saves it from being picked by humans, a fate endangering many of the showier orchid species. Despite its relative success, there is evidence that populations are declining.

Taxonomy

''Neottia'' is

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

for bird's nest, and comes from the appearance of the roots in '' Neottia nidus-avis''.South West Colorado Wildflowers
/ref> The species epithet ''ovata'' comes from the

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

ovatus, meaning ovate or egg-shaped, and describing the leaves of the plant. Twayblade is an old English word meaning two-leaf. The common name in Germany (Zweiblatt) is very similar and has the same meaning. Common twayblade was formerly placed in the genus ''Listera'', but molecular phylogenetic studies have shown that '' Neottia nidus-avis'', the bird's-nest orchid, evolved within the same group,, p. 864 and the two genera have been combined.

References

* CLAESSENS, J. & J. KLEYNEN : The flower of the European Orchid – Form and function, 2011, .

External links

* {{Taxonbar, from=Q15502088 Orchids of Europe Orchids of Asia Plants described in 1753 Taxa named by Carl Linnaeus ovata