''Neosauroides'' is an

ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxon'' comes from the Ancient Greek (''íchnos'') meaning "track" and English , itself derived from ...

of probable

scincid Skinks are a type of lizard belonging to the family (biology), family Scincidae, a family in the Taxonomic rank, infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one o ...

lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...

mainly known from the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

(

Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...

) of

South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...

. The ichnogenus contains two named ichnospecies, ''N. koreanensis'' and ''N. innovatus'', representing the first lizard trackway reported from the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...

period.

Discovery and naming

The ichnogenus ''Neosauroides'' means "new reptile" due to its novelty for being the only known lizard trackway from the Cretaceous period at the time of its initial description, and the type ichnospecies ''N. koreanensis'' from the

Haman Formation The Haman Formation () is an Early Cretaceous geological formation in South Korea. It has been dated to the Albian, with an estimated maximum depositional age of 105.4 ± 0.4 Ma. The deposit is known for its tracks, including those of dinos ...

is named in reference to Korea. The second ichnospecies ''N. innovatus'' from the

Jinju Formation The Jinju Formation (), also known as the Dongmyeong Formation () in some literature is an Early Cretaceous geologic Formation (geology), formation in South Korea.Hekou Group The Hekou Group is a geological group in Gansu Province, China. It is Early Cretaceous in age. Many dinosaur fossils have been recovered from the Hekou Group, including iguanodonts, large sauropods, and Thyreophora, armored dinosaurs. Fossil eggs ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

, and from the

Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...

( Calabrian) Bacons Castle Formation of the United States.

Classification

The trackmaker of ''Neosauroides'' is probably a lizard, but its taxonomic identity is uncertain. The original description of ''N. koreanensis'' did not assign this ichnotaxon to a specific clade, while the describers of ''N. innovatus'' included ''Neosauroides'' within the infraclass

Lepidosauromorpha Lepidosauromorpha (in PhyloCode known as Pan-Lepidosauria) is a group of reptiles comprising all diapsids closer to lizards than to archosaurs (which include crocodiles and birds). The only living sub-group is the Lepidosauria, which contains tw ...

which includes all reptiles closer to lizards than

archosaur Archosauria () or archosaurs () is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only extant taxon, extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistics ...

s such as

crocodilian Crocodilia () is an Order (biology), order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorp ...

s and

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s. The authors also considered the other lizard ichnotaxon from the

Hasandong Formation The Hasandong Formation () is an Early Cretaceous (Aptian to Albian) geologic Formation (geology), formation in South Korea. It has been dated to the late Aptian and earliest Albian, between 118.0 ± 2.6 Ma and 112.4 ± 1.3 Ma. Dinosaur ...

(

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

) of South Korea, ''Sauripes hadongensis'', as a potential ''

nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...

'' due to insufficient description and problematic holotype designation, which would make ''Neosauroides'' to be the only valid Cretaceous lizard ichnotaxon. In 2021, Weems suggested that ''Neosauroides'' represents

skink Skinks are a type of lizard belonging to the family (biology), family Scincidae, a family in the Taxonomic rank, infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one o ...

tracks based on an isolated footprint identified as ''N.'' isp. from the Early Pleistocene of the United States, the only record of this ichnogenus outside Cretaceous East Asia. As of 2024, the oldest known skink ''Electroscincus zedi'' is reported from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

(

Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...

)

Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...

, slightly younger than the

-aged Haman and Jinju Formation.

References

Trace fossils {{trace-fossil-stub