''Navajodactylus'' (meaning "Navajo finger") is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the Order (biology), order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cre ...

from

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

(late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...

stage) deposits of the

San Juan Basin The San Juan Basin is a geologic structural basin located near the Four Corners region of the Southwestern United States. The basin covers 7,500 square miles and resides in northwestern New Mexico, southwestern Colorado, and parts of Utah ...

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...

Discovery

The holotype specimen of ''Navajodactylus'' was discovered and collected by

oceanographer Oceanography (), also known as oceanology and ocean science, is the scientific study of the oceans. It is an Earth science, which covers a wide range of topics, including ecosystem dynamics; ocean currents, waves, and geophysical fluid dynamics ...

Arjan C. Boeré from the

Kirtland Formation The Kirtland Formation (originally the Kirtland Shale) is a sedimentary geological formation. Description The Kirtland Formation is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain t ...

in 2002. ''Navajodactylus'' was first named by Robert M. Sullivan and Denver W. Fowler in 2011 and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

is ''Navajodactylus boerei''. The generic name honors the

Navajo Nation The Navajo Nation ( nv, Naabeehó Bináhásdzo), also known as Navajoland, is a Native Americans in the United States, Native American Indian reservation, reservation in the United States. It occupies portions of northeastern Arizona, northwe ...

, combining their name with a Greek δάκτυλος, ''daktylos'', "finger". The specific name honors Arjan C. Boeré. ''Navajodactylus'' is based on the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

SMP SMP may refer to: Organisations * Scale Model Products, 1950s, acquired by Aluminum Model Toys * School Mathematics Project, UK developer of mathematics textbooks * '' Sekolah Menengah Pertama'', "junior high school" in Indonesia * Shanghai Mun ...

VP-1445, from the Hunter Wash Member of the

, San Juan Basin, dating to the upper Campanian, about 75 million years old. It consists of three pieces of the first phalanx of the wing finger. The

paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...

is SMP VP-1853, an ulna fragment. Two first phalanges,

TMP TMP can refer to any of the following: Chemistry * 2,2,6,6-Tetramethylpiperidine, an organic chemistry reagent * Thymidine monophosphate, a nucleotide * Trimethoprim, an antibiotic * Trimethyl phosphate, a solvent * Trimethylolpropane, a precurso ...

72.1.1 and TMP 82.19.295, both from the

Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...

of the Dinosaur Provincial Park, Alberta, were referred to this taxon, but in 2019, Hone ''et al.'' recognized these as referable to their new azhdarchid '' Cryodrakon''.

Description

''Navajodactylus'' was a medium-sized pterosaur, with an estimated

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...

of 3.5 meters (11.5 ft). Its

autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

largely exist in the unique form of the process on the first wing phalanx for the extensor tendon.

Systematics

''Navajodactylus'' was tentatively assigned to the family

Azhdarchidae Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cre ...

because of its geological age, however, it does not show any

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...

of the group. Indeed, it may not actually be an azhdarchid, as it lacks pneumacy in its forelimb elements, which is opposed to the extensive pneumacy seen in azhdarchids.Wilton, Mark P. (2013). Pterosaurs: Natural History, Evolution, Anatomy.

Princeton University Press Princeton University Press is an independent Academic publishing, publisher with close connections to Princeton University. Its mission is to disseminate scholarship within academia and society at large. The press was founded by Whitney Darrow, ...

. .

References

{{Portal bar, Paleontology, Cretaceous, United States Azhdarchids Late Cretaceous pterosaurs of North America Fossil taxa described in 2011

Discovery

Description

Systematics

See also

References