Nannoperca Australis
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The southern pygmy perch (''Nannoperca australis''), also known as the Tasmanian pygmy perch, is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
, a
temperate perch Percichthyidae, the temperate perches, are a family of freshwater ray-finned fish found in Australia and southern South America. Formerly placed in the order Perciformes, they are now placed in the order Centrarchiformes. The name Percichthyida ...
from the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Percichthyidae which is native to south-eastern Australia and
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
.


Description

The southern pygmy perch has an oblong and moderately compressed body with a convex dorsal profile and a straight ventral profile. It has a large head the top of which bulges slightly and a blunt snout. It has a slightly oblique, terminal mouth which is protractile with the
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
reaching to a level near the centre of the eye. There are thin bands of villiform teeth on the jaws and the roof of the mouth. There is a
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...
made up of an irregular series of pored scales. The dorsal fin has 7-9 spines in its front part, separated from the rear part by a deep notch, the rear part contains 7-10 soft rays. The
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
is similar in shape to the soft part of the
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
and has 3 spines and 7-8 soft rays. The large
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
is rounded. They are variable in colour, the colour apparently being determined by their environment and can be pale cream through to greenish-brown, with paler underparts. They can also show irregular markings on the flanks such as dark spotting or horizontal stripes. In the breeding season the males are brighter as their dorsal, anal and caudal fins develop a red colour and have black margins and the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...
s and the area around the anus become black. These fish attain a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of but are more commonly recorded at standard lengths of around .


Distribution

Southern pygmy perch are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to temperate rivers systems in southeastern Australia, especially the
Murray Murray may refer to: Businesses * Murray (bicycle company), an American bicycle manufacturer * Murray Motor Car Company, an American car manufacturer * Murrays, an Australian bus company * Murray International Trust, a Scottish investment trus ...
- Murrumbidgee river system from Brinagagee near
Darlington Point Darlington Point is a small town on the banks of the Murrumbidgee River in the central Riverina district of western New South Wales, Australia. It is part of the Murrumbidgee Council local government area.  The centre of town is four kilomet ...
in
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
, in
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
, west to Ewens Ponds in
South Australia South Australia (commonly abbreviated as SA) is a States and territories of Australia, state in the southern central part of Australia. With a total land area of , it is the fourth-largest of Australia's states and territories by area, which in ...
. They also occur on
Flinders Island Flinders Island, the largest island in the Furneaux Group, is a island in the Bass Strait, northeast of the island of Tasmania. Today Flinders Island is part of the state of Tasmania, Australia. It is from Cape Portland, Tasmania, Cape Portl ...
and on King Island in the
Bass Strait Bass Strait () is a strait separating the island state of Tasmania from the Mainland Australia, Australian mainland (more specifically the coast of Victoria (Australia), Victoria, with the exception of the land border across Boundary Islet). The ...
and in the northward draining rivers in Tasmania.


Habitat and biology

Southern pygmy perch are found in a wide variety of
freshwater habitat Freshwater ecosystems are a subset of Earth's aquatic ecosystems that include the biological communities inhabiting freshwater waterbodies such as lakes, ponds, rivers, streams, springs, bogs, and wetlands. They can be contrasted with marine ec ...
s so long as they have
aquatic vegetation Aquatic plants, also referred to as hydrophytes, are vascular plants and Non-vascular plant, non-vascular plants that have adapted to live in aquatic ecosystem, aquatic environments (marine ecosystem, saltwater or freshwater ecosystem, freshwater ...
. They can be found in both still or slow flowing waters and they have been recorded in low current streams, lakes,
billabong In Australian English, a billabong ( ) is a small body of water, usually permanent. It is usually an oxbow lake caused by a change in course of a river or creek, but other types of small lakes, ponds or waterholes are also called billabongs ...
s, ditches, impoundments,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and
ephemeral Ephemerality (from the Greek word , meaning 'lasting only one day') is the concept of things being transitory, existing only briefly. Academically, the term ephemeral constitutionally describes a diverse assortment of things and experiences, fr ...
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s. In all of these habitat they prefer the vegetated marginal areas. It is a mainly
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
species with its diet mostly consisting of small
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s such as
mosquito larva Mosquitoes, the Culicidae, are a family of small flies consisting of 3,600 species. The word ''mosquito'' (formed by '' mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mosquitoes have a slender segmented body, ...
e and other
aquatic insect Aquatic insects or water insects live some portion of their life cycle in the water. They feed in the same ways as other insects. Some ''diving'' insects, such as predatory diving beetles, can hunt for food underwater where land-living insects ...
s, as well as ''
Daphnia ''Daphnia'' is a genus of small planktonic crustaceans, in length. ''Daphnia'' are members of the Order (biology), order Anomopoda, and are one of the several small aquatic crustaceans commonly called water fleas because their Saltation (gait), ...
'' and other
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s. As they mature they prey on larger insects including
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...
but their small mouth means that they do not consume many other fishes. Southern pygmy perch attain
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
during the first year after hatching, mature males are mature at about and females at around . The breeding season runs from the late winter into the early spring once the water temperatures rise above . Breeding males are
territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
and defend a small area of water from other males. The female lays small clutches of eggs in the aquatic vegetation or onto the riverbed within the male's territory. These hatch into larvae of 3-4mm in length 2-4 days after fertilisation.


Conservation


Status

The Southern pygmy perch was formerly widespread in New South Wales, but has now largely been
extirpated Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...
from the state. there were only three known locations where it was still found: in the
Lachlan River The Lachlan River (Wiradjuri: ''Kalari'', ''Galiyarr'') is an intermittent river that is part of the Murrumbidgee catchment within the Murray–Darling basin, located in the Southern Tablelands, Central West, and Riverina regions of New Sou ...
catchment, north of Yass; as near Holbrook, and
Albury Albury (; ) is a major regional city that is located in the Murray River, Murray region of New South Wales, Australia. It is part of the twin city of Albury–Wodonga, Albury-Wodonga and is located on the Hume Highway and the northern side of ...
(
Tumbarumba Tumbarumba ( ) is a town in New South Wales, Australia, about southwest of the state capital, Sydney. Tumbarumba is located in the South West Slopes (New South Wales), South West Slopes region at the western edge of the Snowy Mountains. The s ...
?). In 2021 a fourth population was found in the Ooolong Creek in the town of
Dalton Dalton may refer to: Science * Dalton (crater), a lunar crater * Dalton (program), chemistry software * Dalton (unit) (Da), a.k.a. unified atomic mass unit * John Dalton, chemist, physicist and meteorologist * 12292 Dalton, an asteroid Ent ...
, by two teenagers. As of 2007 the fish was still common in coastal Victoria, but rare in the
Murray River The Murray River (in South Australia: River Murray; Ngarrindjeri language, Ngarrindjeri: ''Millewa'', Yorta Yorta language, Yorta Yorta: ''Dhungala'' or ''Tongala'') is a river in Southeastern Australia. It is List of rivers of Australia, Aust ...
catchment of that state, and rare and threatened in South Australia. In Tasmania it remains relatively common. The reasons for the decline and continuing threats to this species includes invasive alien fish species such as
eastern mosquitofish The eastern mosquitofish (''Gambusia holbrooki'') is a species of freshwater fish, closely related to the western mosquitofish, ''Gambusia affinis''. It is a member of the family Poeciliidae of order Cyprinodontiformes. The eastern mosquitofish i ...
(which feed on their eggs and
compete Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, individ ...
for food and territories),
brown trout The brown trout (''Salmo trutta'') is a species of salmonid ray-finned fish and the most widely distributed species of the genus ''Salmo'', endemic to most of Europe, West Asia and parts of North Africa, and has been widely introduced globally ...
,
Eurasian carp The common carp (''Cyprinus carpio''), also known as European carp, Eurasian carp, or simply carp, is a widespread freshwater fish of eutrophic waters in lakes and large rivers in Europe and Asia.Fishbase''Cyprinus carpio'' Linnaeus, 1758/ref>Ark ...
and
European perch The European perch (''Perca fluviatilis''), also known as the common perch, redfin perch, big-scaled redfin, English perch, Euro perch, Eurasian perch, Eurasian river perch, Hatch, poor man's rockfish or in Anglophone parts of Europe, simply the ...
("redfins"), as well as cold water pollution, and
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological proces ...
and
destruction Destruction may refer to: Concepts * Destruktion, a term from the philosophy of Martin Heidegger * Destructive narcissism, a pathological form of narcissism * Self-destructive behaviour, a widely used phrase that ''conceptualises'' certain kin ...
. The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
assessed the conservation status of this species as
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
in 2019.


Measures

The Department of Primary Industries in New South Wales has undertaken a
captive breeding Captive breeding, also known as captive propagation, is the process of keeping plants or animals in controlled environments, such as wildlife reserves, zoos, Botanical garden, botanic gardens, and other Conservation biology, conservation facilitie ...
programme for this species. In August 2023, a group of volunteers working with OzFish Unlimited and the local council undertook the translocation of 200 fish to Oolong Creek, to boost the population there.


Taxonomy

The Southern pygmy perch was first formally described in 1861 by Albert Günther with the type locality given as the Murray River. Molecular studies have shown that there are two cryptic species within ''Nannoperca australis'' one from the eastern coastal streams and the other from the
Murray–Darling basin The Murray–Darling Basin is a large geographical area in the interior of southeastern Australia, encompassing the drainage basin of the tributaries of the Murray River, Australia's longest river, and the Darling River, a right tributary of ...
and the western coastal streams.


Aquarium

The Southern pygmy perch is in the
aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...
trade within Australia, where it is kept in cold water aquaria and in ponds.


References


External links


Southern Pygmy Perch
{{Taxonbar, from=Q5747401 Nannoperca Freshwater fish of Australia Fish described in 1861 Taxa named by Albert Günther