Myrsinoideae is a subfamily of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Primulaceae The Primulaceae ( ), commonly known as the primrose family (but not related to the Onagraceae, evening primrose family), are a family (biology), family of Herbaceous plant, herbaceous and woody flowering plants including some favourite garden pla ...

in the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Ericales The Ericales are a large and diverse order of flowering plants in the asterid group of the eudicots. Well-known and economically important members of this order include tea and ornamental camellias, persimmon, ebony, blueberry, cranberry, l ...

. It was formerly recognized as the family Myrsinaceae, or the myrsine family, consisting of 35 genera and about 1000 species. It is widespread in temperate to tropical climates extending north to

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

, and

Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...

, and south to

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

, and

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

. Plants are mostly

mesophytic Mesophytes are terrestrial plants which are adapted to neither particularly dry nor particularly wet environments. An example of a mesophytic habitat would be a rural temperate meadow, which might contain goldenrod, clover, oxeye daisy, and '' Ros ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

s and

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

s; a few are

liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...

s or subherbaceous. Their leathery,

evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are

simple Simple or SIMPLE may refer to: *Simplicity, the state or quality of being simple Arts and entertainment * ''Simple'' (album), by Andy Yorke, 2008, and its title track * "Simple" (Florida Georgia Line song), 2018 * "Simple", a song by John ...

and

alternate Alternative or alternate may refer to: Arts, entertainment and media * Alternative (''Kamen Rider''), a character in the Japanese TV series ''Kamen Rider Ryuki'' * Alternative comics, or independent comics are an alternative to mainstream superh ...

, with smooth margins and without

stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole (botany), petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part ...

s. They are often dotted with glands and resinous cavities. The latter may take the form of secretory lines. The plants are mostly

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...

, but a few are

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

. Their small

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s are arranged in

racemose A raceme () or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the shoo ...

terminal clusters, or in the leaf

axils A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ...

. The flowers have four or five

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s and

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s. The floral envelope (

perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...

) has a distinct calyx and corolla. The calyx is regular and polysepalous. The nonfleshy petals of the corolla are more or less united, closely overlapping. The four or five

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s are usually isomerous with the perianth. The

carpel Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more ...

has one

style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...

and one stigma, with the

ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...

unilocular, superior or semi-inferior. The one-seeded,

indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part. Structures that ...

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is a thin-fleshed

berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...

drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...

. North American species are the marlberry (''Ardisia escalloniodes'') and the Florida rapanea (''Myrsine cubana''). Plants in the subfamily have few economic uses. A few genera, such as '' Ardisia'', '' Cyclamen'', ''

Lysimachia ''Lysimachia'' ( ) is a genus consisting of 182 accepted species of flowering plants traditionally classified in the family Primulaceae. Based on a molecular phylogenetic study it was transferred to the family Myrsinaceae, before this family w ...

'', and ''

Myrsine ''Myrsine'' is a genus of flowering plants in the family Primulaceae. It was formerly placed in the family Myrsinaceae before this was merged into the Primulaceae. It is found nearly worldwide, primarily in tropical and subtropical areas. It co ...

'', are grown as

ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

s, especially ''Ardisia crispa'' and ''Myrsine africana''. One species, '' Ardisia japonica'' (Chinese: 紫金牛; pinyin: ''zǐjīn niú''), is one of the 50 fundamental herbs in

traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medicine, alternative medical practice drawn from traditional medicine in China. A large share of its claims are pseudoscientific, with the majority of treatments having no robust evidence ...

. In the

APG III system The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a f ...

and onwards, the Myrsinaceae were not recognized, but were sunk into

, which in that system is circumscribed very broadly.

Genera

* '' Aegiceras'' * '' Amblyanthopsis'' * '' Amblyanthus'' * ''

Anagallis ''Anagallis'' is a formerly recognized genus of flowering plants in the family (biology), family Primulaceae. It had about 20–25 species, commonly called pimpernels. Molecular phylogenetics, Molecular phylogenetic studies showed that ''Anagal ...

'' * ''

Antistrophe Antistrophe (, "a turning back") is the portion of an ode sung by the chorus in its returning movement from west to east in response to the strophe, which was sung from east to west. Characteristics Usage as a literary device It has the n ...

'' * '' Ardisia'' * '' Asterolinon'' (should be included into the ''Anagallis'' clade) * '' Badula'' * '' Conandrium'' * '' Coris'' * '' Ctenardisia'' * '' Cybianthus'' * '' Cyclamen'' * '' Discocalyx'' * '' Elingamita'' * '' Embelia'' * '' Emblemantha'' * '' Fittingia'' * '' Geissanthus'' * '' Glaux'' (should be included into the clade'' Lysimachia'') * '' Heberdenia'' * '' Hymenandra'' * '' Labisia'' * '' Loheria'' * ''

'' * '' Monoporus'' * ''

'' * '' Oncostemum'' * '' Parathesis'' * '' Pelletiera'' (should be included into the'' Anagallis clade'') * '' Pleiomeris'' * '' Rapanea'' * '' Sadiria'' * '' Solonia'' * '' Stylogyne'' * '' Tapeinosperma'' * '' Trientalis'' * '' Tetrardisia'' * '' Vegaea'' * '' Wallenia'' The following genera, traditionally categorized in

''sensu lato'', should, according to Källersjö et al. (2000), belong to the Myrsinoideae (the clade of Myrsinaceae ''s. l.''): ''Anagallis'', ''Ardisiandra'', ''Asterolinon'', ''Coris'', ''Cyclamen'', ''Glaux'', ''Lysimachia'', ''Pelletiera '' and ''Trientalis''.

References

* * * {{Authority control Asterid subfamilies Primulaceae