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''Mycalesis'', the bushbrowns, are a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of brush-footed
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises t ...
. They are common in the warm regions from
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the former ...
to Australia, and have a high diversity in
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
and the
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as ...
. They are notably
polymorphic Polymorphism, polymorphic, polymorph, polymorphous, or polymorphy may refer to: Computing * Polymorphism (computer science), the ability in programming to present the same programming interface for differing underlying forms * Ad hoc polymorphism ...
, with wet- and dry-season forms differing in many species, especially as regards size and number of underwing eyespots. ''Mycalesis'' superficially resemble the species ''
Orsotriaena medus ''Orsotriaena medus'' is a butterfly found in south Asia, southeast Asia, and Australia. It is the only species in the genus ''Orsotriaena'', first described by Hans Daniel Johan Wallengren in 1858. Historical name The butterfly has historic ...
'', but can readily be identified by the number of spots.


Species

Listed alphabetically: * '' Mycalesis adamsoni'' Watson, 1897 – Watson's bushbrown * ''
Mycalesis adolphei ''Heteropsis adolphei'', the red-eye bushbrown, is a species of satyrine butterfly found in southern India. The species name is after Adolphe Delessert who collected the first specimens based on which the species was described. Description Uppe ...
'' (Guérin-Ménéville, 1843) – redeye bushbrown * ''
Mycalesis aethiops ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Butler, 1868 * '' Mycalesis amoena'' Druce, 1873 * ''
Mycalesis anaxioides ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Marshall & de Nicéville, 1883 * ''
Mycalesis annamitica ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to ...
'' Fruhstorfer, 1906 – Annam bushbrown, tawny bush-brown * ''
Mycalesis anapita ''Mycalesis anapita'', the tawny bush-brown, is a species of Satyrinae butterfly described by Frederic Moore in 1858. It is found in southern Burma, southern Thailand, Peninsular Malaysia, Sumatra and Borneo.Kirton, L. 2001, ''A Naturalists Guid ...
'' Moore,
858 __NOTOC__ Year 858 ( DCCCLVIII) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – King Louis the German, summoned by the disaffected Frankish ...
/small> * '' Mycalesis anaxias'' Hewitson, 1862 – white-bar bushbrown * ''
Mycalesis arabella ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Fruhstorfer, 1906 * ''
Mycalesis aramis ''Mycalesis aramis'' is a butterfly of the family Nymphalidae. It is found on Luzon in the Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * ...
'' Hewitson, 1866 * ''
Mycalesis asophis ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Hewitson, 1862 * ''
Mycalesis atrata ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Röber, 1887 * ''
Mycalesis barbara ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Grose-Smith, 1894 * ''
Mycalesis bazochii ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' (Guérin-Méneville, 1831) * ''
Mycalesis biformis ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Rothschild, 1916 * ''
Mycalesis bilineata ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Fruhstorfer, 1906 * ''
Mycalesis bisaya ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' C. & R. Felder, 1863 * ''
Mycalesis cacodaemon ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Kirsch, 1877 * ''
Mycalesis comes ''Mycalesis'', the bushbrowns, are a genus of Nymphalidae, brush-footed butterfly, butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymo ...
'' Grose-Smith, 1894 * ''
Mycalesis deficiens ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Fruhstorfer, 1906 * '' Mycalesis discobolus'' Fruhstorfer, 1906 * '' Mycalesis dohertyi'' Elwes, 1891 * '' Mycalesis drusillodes'' (Oberthür, 1894) * '' Mycalesis duponcheli'' (Guérin-Méneville, 1830) * '' Mycalesis durga'' Grose-Smith & Kirby, 1896 * '' Mycalesis elia'' Grose-Smith, 1894 * '' Mycalesis evansii'' Tytler, 1914 – Tytler's bushbrown * ''
Mycalesis evara ''Mycalesis'', the bushbrowns, are a genus of Nymphalidae, brush-footed butterfly, butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymo ...
'' Fruhstorfer, 1906 * '' Mycalesis felderi'' Butler, 1868 * ''
Mycalesis francisca ''Mycalesis francisca'' is an East Palearctic species of satyrine butterfly found in China, Japan, Assam, Burma and Indochina The larva feeds on ''Imperata cylindrica'', ''Miscanthus sinensis'', ''Setaria palmifolia'' and ''Oplismenus undulatif ...
'' (Stoll,
780 __NOTOC__ Year 780 ( DCCLXXX) was a leap year starting on Saturday of the Julian calendar. The denomination 780 for this year has been used since the early medieval period, when the Anno Domini calendar era became the prevalent method in Eur ...
– lilacine bushbrown ** ''Mycalesis francisca formosana'' Fruhstorfer, 1908 * '' Mycalesis fuscum'' (C. & R. Felder, 1860) – Malayan bush-brown * '' Mycalesis giamana'' Parsons, 1986 * ''
Mycalesis gotama ''Mycalesis gotama'', the Chinese bushbrown, is an East Palearctic species of satyrine butterfly found in China, Japan, Assam, Burma, the northeastern Himalayas and northern Vietnam. The larva feeds on ''Miscanthus sinensis'', ''Miscanthus fl ...
'' Moore, 1857 – Chinese bushbrown ** ''Mycalesis gotama nanda'' Fruhstorfer, 1908 * ''
Mycalesis heri ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Moore, 1857 – Moore's bushbrown * '' Mycalesis horsfieldi'' (Moore,
892 Year 892 ( DCCCXCII) was a leap year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – Poppo II, duke of Thuringia (Central Germany), is deposed by King Arnu ...
Malaysia, Java, Celebes. See
aps APS or Aps or aps or similar may refer to: Education * Abbottabad Public School * Adarsh Public School, a public school in New Delhi, India * Alamogordo Public Schools * Albuquerque Public Schools, New Mexico, US school district * Allendale Publ ...
– Horsfield's bush brown * ''
Mycalesis igilia ''Mycalesis igilia'', the small long-brand bushbrown, is a species of Satyrinae, satyrid butterfly found in south India.Mycalesis igoleta'' C. & R. Felder, 1863 * ''
Mycalesis inayoshii ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Aoki & Yamaguchi, 1995 * '' Mycalesis inopia'' Fruhstorfer, 1908 * ''
Mycalesis intermedia ''Mycalesis intermedia'', the intermediate bushbrown, is a species of satyrine butterfly found in Burma, Cambodia, Thailand, Peninsular Malaysia, Langkawi Island, Indochina, southern Yunnan and India India, officially the Republic of In ...
'' (Moore,
892 Year 892 ( DCCCXCII) was a leap year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – Poppo II, duke of Thuringia (Central Germany), is deposed by King Arnu ...
– intermediate bushbrown * ''
Mycalesis ita ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' C. & R. Felder, 1863 * ''
Mycalesis itys ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' C. & R. Felder,
867 __NOTOC__ Year 867 ( DCCCLXVII) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * September 23 – Emperor Michael III is murdered, by order o ...
/small> – Itys bush brown * '' Mycalesis janardana'' Moore, 1857 – Common Bush Brown, Janardana Bush Brown, Mottled Bush-Brown * '' Mycalesis khasia'' Evans, 1920 – palebrand bushbrown * ''
Mycalesis lepcha ''Mycalesis'' (''annamitica'') ''lepcha'', the Lepcha bushbrown, is a satyrine butterfly found in Asia. It is not resolved whether it is best considered a distinct species, or included in ''Mycalesis annamitica''. It was formerly included in ''M ...
'' (Moore, 1880) – Lepcha bushbrown * ''
Mycalesis madjicosa ''Mycalesis madjicosa'' is a satyrine butterfly endemic to Japan. The larva feeds on ''Imperata'' and ''Cortaderia selloana''. Subspecies *''M. m. madjicosa'' *''M. m. amamiana'' Fujioka, 1975 (Amami, Okinawa) References External linksImage ...
'' Butler, 1868 * '' Mycalesis maianeas'' Hewitson, 1864 * '' Mycalesis malsarida'' Butler, 1868 – plain bushbrown * ''
Mycalesis malsara ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and d ...
'' Moore, 1857 – white-line bushbrown * '' Mycalesis mamerta'' (Stoll,
780 __NOTOC__ Year 780 ( DCCLXXX) was a leap year starting on Saturday of the Julian calendar. The denomination 780 for this year has been used since the early medieval period, when the Anno Domini calendar era became the prevalent method in Eur ...
– blind-eye bushbrown ** '' Mycalesis mamerata davisoni'' – Palni bushbrown * '' Mycalesis manii'' Doherty, 1886 * '' Mycalesis maura'' Grose-Smith, 1894 * '' Mycalesis mnasicles'' Hewitson, 1864 * '' Mycalesis meeki'' Rothschild, 1915 * '' Mycalesis mehadeva'' (Boisduval, 1832) * ''
Mycalesis mercea ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Evans (''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate des ...
''?) – Pachmarhi bushbrown * '' Mycalesis messene'' Hewitson, 1862 * '' Mycalesis mestra'' Hewitson, 1862 – white-edged bushbrown * ''
Mycalesis mineus ''Mycalesis mineus'', the dark-brand bush brown,
Butterflies of Singapore is a
'' (Linnaeus, 1758) – dark-branded bushbrownKirton, Laurence G. (2014). ''A Naturalist's Guide to the Butterflies of Peninsular Malayasia, Singapore and Thailand.'' Oxford:John Beaufoy Publ., p.62. * '' Mycalesis misenus'' de Nicéville, 1889 – salmon-branded bushbrown * '' Mycalesis mnasicles'' Hewitson, 1864 * '' Mycalesis moorei'' C. & R. Felder,
867 __NOTOC__ Year 867 ( DCCCLXVII) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * September 23 – Emperor Michael III is murdered, by order o ...
/small> * '' Mycalesis mucia'' Hewitson, 1862 * '' Mycalesis mulleri'' Tennent, 2000 * '' Mycalesis mynois'' Hewitson, 1864 * '' Mycalesis mystes'' de Nicéville, 1891 - many-tufted bushbrown * ''
Mycalesis nala ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' C. & R. Felder, 1859 * '' Mycalesis nerida'' Grose-Smith, 1902 * '' Mycalesis nicotia'' Westwood,
850 ''For codepage, see CP850.'' __NOTOC__ Year 850 (Roman numerals, DCCCL) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Europe * February 1 – King Ramiro I of A ...
/small> – bright-eye bushbrown * ''
Mycalesis oculus ''Mycalesis oculus'', the red-disc bushbrown, is a satyrine butterfly found in southern India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most ...
'' Marshall, 1880 – red-disc bushbrown * ''
Mycalesis oroatis ''Mycalesis oroatis'' is a butterfly of the family Nymphalidae. It is found in South-east Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geograph ...
'' Hewitson, 1864 * ''
Mycalesis orseis ''Mycalesis orseis'', the purple bushbrown, is a butterfly of the family Nymphalidae. It is found in South-east Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia o ...
'' Hewitson, 1864 – purple bushbrown * ''
Mycalesis panthaka ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Fruhstorfer, 1909 * '' Mycalesis patiana'' Eliot, 1969 * '' Mycalesis pitana'' Staudinger,
897 __NOTOC__ Year 897 ( DCCCXCVII) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Spring – King Lambert II travels to Rome with his mother, Queen Agelt ...
/small> * ''
Mycalesis patnia ''Mycalesis patnia'', the glad-eye bushbrown, is a Satyrinae, satyrid butterfly found in southern India and Sri Lanka. Description Upperside: The male has a dark umber-brown ground colour slightly suffused with ochraceous. Forewings and hindw ...
'' Moore, 1857 – glad-eye bushbrown * ''
Mycalesis perseoides ''Mycalesis perseoides'', the Pachmarhi bushbrown, is a species of satyrine butterfly (family Nymphalidae) found in Asia (Burma, Thailand, Langkawi Island, Peninsular Malaya, Indochina, Yunnan, India) Description This form closely resembles '' ...
'' (Moore,
892 Year 892 ( DCCCXCII) was a leap year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – Poppo II, duke of Thuringia (Central Germany), is deposed by King Arnu ...
– Pachmarhi bushbrown * ''
Mycalesis perseus ''Mycalesis perseus'', the dingy bushbrown or common bushbrown, is a species of satyrine butterfly found in south Asia and southeast Asia. Description Wet-season form. Male and female. Upperside dark to somewhat pale vandyke-brown. Fore wing w ...
'' (Fabricius, 1775) – dingy bushbrown or common bushbrown ** ''Mycalesis perseus blasius'' (Fabricius, 1798) * '' Mycalesis pitana'' Staudinger,
897 __NOTOC__ Year 897 ( DCCCXCVII) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Spring – King Lambert II travels to Rome with his mother, Queen Agelt ...
/small> * '' Mycalesis phidon'' Hewitson, 1862 * ''
Mycalesis radza ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Moore, 1878 * ''
Mycalesis rama ''Mycalesis rama'', the Cingalese bushbrown, is a species of satyrine butterfly of family Nymphalidae. It is a very shy animal, distributed in bamboo forests. It is endemic to Sri Lanka. Wingspan is about 56mm. Dorsal surface is dark brown wit ...
'' (Moore, 1892) – Cingalese bushbrown * '' Mycalesis sangaica'' Butler, 1877 – single ring bushbrown ** ''Mycalesis sangaica mara'' Fruhstorfer, 1908 * '' Mycalesis shiva'' (Boisduval, 1832) * '' Mycalesis siamica'' Riley & Godfrey, 1921 * ''
Mycalesis sirius ''Mycalesis'', the bushbrowns, are a genus of Nymphalidae, brush-footed butterfly, butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymo ...
'' (Fabricius, 1775) * '' Mycalesis splendens'' Mathew, 1887 * ''
Mycalesis suaveolens ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Wood-Mason & de Nicéville, 1883 – Wood-Mason's bushbrown ** ''Mycalesis suaveolens kagina'' Fruhstorfer, 1908 * '' Mycalesis subpersa'' Rothschild, 1915 * ''
Mycalesis sudra ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' C. & R. Felder,
867 __NOTOC__ Year 867 ( DCCCLXVII) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * September 23 – Emperor Michael III is murdered, by order o ...
/small> * '' Mycalesis tagala'' C. & R. Felder, 1863 * '' Mycalesis terminus'' (Fabricius, 1775) – orange bushbrown * ''
Mycalesis thailandica ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Aoki & Yamaguchi, 1984 * '' Mycalesis thyateira'' (Fruhstorfer, 1911) * ''
Mycalesis treadawayi ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and dr ...
'' Schröder, 1976 * '' Mycalesis unica'' Leech,
892 Year 892 ( DCCCXCII) was a leap year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – Poppo II, duke of Thuringia (Central Germany), is deposed by King Arnu ...
/small> * '' Mycalesis valeria'' Grose-Smith, 1900 * ''
Mycalesis valeriana ''Mycalesis'', the bushbrowns, are a genus of Nymphalidae, brush-footed butterfly, butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymo ...
'' Grose-Smith, 1900 * ''
Mycalesis visala ''Mycalesis visala'', the long-brand bushbrown, is a species of satyrine butterfly found in south Asia. It might include the Tamil bushbrown as a subspecies. Description The colour and wing markings of both seasonal forms of ''M. visala'' cl ...
'' Moore,
858 __NOTOC__ Year 858 ( DCCCLVIII) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Summer – King Louis the German, summoned by the disaffected Frankish ...
/small> – long-brand bushbrown ** '' Mycalesis visala subdita'' – Tamil bushbrown * '' Mycalesis wayewa'' Doherty, 1891 * '' Mycalesis zonata'' Matsumura, 1909 – South China bushbrown


See also

*''
Orsotriaena ''Orsotriaena medus'' is a butterfly found in south Asia, southeast Asia, and Australia. It is the only species in the genus ''Orsotriaena'', first described by Hans Daniel Johan Wallengren in 1858. Historical name The butterfly has historic ...
''


References

{{Taxonbar, from=Q6946839 Satyrini Nymphalidae genera Taxa named by Jacob Hübner