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The mutton snapper (''Lutjanus analis'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
, a snapper belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Lutjanidae Lutjanidae or snappers are a family of perciform fishes, mainly marine but with some members inhabiting estuaries and, in some cases, fresh water (e.g., '' Lutjanus goldiei''). The family includes about 113 species. Most species are used for foo ...
. It is found in the Western Atlantic Ocean.


Taxonomy

The mutton snapper was first formally described in 1828 by the
French French may refer to: * Something of, from, or related to France ** French language, which originated in France ** French people, a nation and ethnic group ** French cuisine, cooking traditions and practices Arts and media * The French (band), ...
zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
Georges Cuvier with the type locality given as Santo Domingo in the Dominican Republic. The specific name refers to the red, sharply pointed anal fin.


Description

The mutton snapper has a moderately deep, almond shaped body, with a nearly lunate
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
. The
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
is almost two lobed and it has a sharply pointed
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
. The
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
is long, extending just past the origin of the anal fin. Both pairs of nostrils are simple holes. It has small eyes, a straight dorsal profile on the forehead and comparatively large terminal mouth with a protrusible upper jaw which is mostly covered by the cheek bone when the mouth is closed. The jaws are equipped with 1-2 tows of conical teeth with some of the outer row being enlarged into canine-like teeth. The vomerine teeth are arranged in a chevron shaped patch with another patch of teeth on each side of the palate. The dorsal fin has 10-11 spines and 13-14 soft rays while the anal fin contains 3 spines and 7-8 soft rays. The pectoral fin has 15-17, typically 16, rays. This is a colourful species with olive green on the back and upper flanks shading to reddish the lower flanks and abdomen. There is a black spot on the between the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...
and the spiny part of the dorsal fin. There is a pair of blue stripes beneath the eyes and the higher one continues on to the operculum. The caudal fin has a black margin. When the fish is resting it has 10 to 12 dark vertical bars on the body, these merge into a solid colour when the fish is swimming. The fins are red. Small juveniles, around in length, have transparent fins and yellowish brown bands on the body, larger juveniles are marked with yellow bands on their body and a more prominent black spot than the adults. This fish attains a maximum total length of , although is more typical, and the maximum published weight is .


Distribution and habitat

The mutton snapper is native to the western Atlantic Ocean where it occurs from
Massachusetts Massachusetts ( ; ), officially the Commonwealth of Massachusetts, is a U.S. state, state in the New England region of the Northeastern United States. It borders the Atlantic Ocean and the Gulf of Maine to its east, Connecticut and Rhode ...
in the north to Brazil in the south. It is commonest in the warmer waters of
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
, the Bahamas, the
Caribbean Sea The Caribbean Sea is a sea of the Atlantic Ocean, North Atlantic Ocean in the tropics of the Western Hemisphere, located south of the Gulf of Mexico and southwest of the Sargasso Sea. It is bounded by the Greater Antilles to the north from Cuba ...
and in the
Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...
. This species occurs in comparatively shallow clear waters at depths from where the water temperature is between . The larger adults live in and around offshore reefs and in area of rock rubble, and the juveniles stay in inshore waters where they can be abundant in habitats such as tidal mangrove creeks, canals and shallow protected bays. The juveniles often shelter within turtle grass beds.


Biology

The mutton snapper is a predatory species which changes its diet during its life cycle. The larvae feed on plankton near the surface of the sea, when they settle to live in the turtle grass beds they eat larger items of plankton and small invertebrates. As they grow they begin to feed on
crustaceans Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of Arthropod, arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquat ...
,
gastropods Gastropods (), commonly known as slugs and snails, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and from the land. Ther ...
and smaller fishes, like grunts and mullets. They feed throughout the day, picking at prey. The body colour changes depending on the method of feeding, the dark barred pattern being adopted when feeding near the bottom. The mutton snapper spawns everywhere it is found, although the main spawning area is in the northeastern Caribbean. They gather in large, transient aggregations to spawn. Spawning takes place in February in the Caribbean while in other parts of its range it occurs in the summer. This species spawns at the same place and at the same time relative to the lunar calendar every year. The females release pelagic eggs that are carried by currents. A female can lay as many as 1,400,000 eggs which the males fertilise externally and which hatch 20 hours after fertilisation. Following hatching, the larvae migrate to shallow water environments within 48 hours. When spawning is finished, the adults move to deeper, offshore waters and become sedentary. The lifespan can be as much as forty years.


Fisheries and conservation

The mutton snapper is an important target species for recreational, commercial and artisanal fisheries. The flesh is considered to be of very high quality and commands high prices at markets throughout its range, especially the meat from the cheek and throat of the larger snappers which are regarded as gourmet foods. This species is caught using hook and line, seines, gill nets, bottom longlines and traps, it is also taken by
spearfishing Spearfishing is fishing using handheld elongated, sharp-pointed tools such as a spear, gig, or harpoon, to impale the fish in the body. It was one of the earliest fishing techniques used by mankind, and has been deployed in artisanal fishi ...
. In Florida it is a very popular with the recreational fisheries, most landing at
Cape Canaveral Cape Canaveral () is a cape (geography), cape in Brevard County, Florida, in the United States, near the center of the state's Atlantic Ocean, Atlantic coast. Officially Cape Kennedy from 1963 to 1973, it lies east of Merritt Island, separated ...
. This species forms predictable spawning aggregations rendering it vulnerable to overfishing while the juveniles are also fished for in some areas worsening the effects of overfishing. In the northern part of its range, stock assessments have shown that there has been a decline of over 60% in commercial landings but the stock is not overfished, there has been a reduction in effort. In Cuba, the size of the known spawning aggregation has decreased. Elsewhere, the mutton snapper is subjected to heavy exploitation and considered to be subject to overfishing in parts of Brazil and Colombia. From the information available it is estimated that the biomass of the stock has declined by at least 20% over a period of 30 to 60 years and so it is listed as
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
by the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
. It is thought that if current trends continue then it should be expected that the status could worsen to vulnerable. The IUCN has recommended the implementation of precautionary management measures, such as the collection of appropriate fisheries data to gain a clearer understanding of the population structures, as well as the protecting important spawning aggregations. Mutton-snapper-lutjanus-analis.jpg Mutton-snapper-eye-closeup.jpg, Eye Mutton-snapper-face-picture.jpg, Face


References


External links

*
Mutton Snapper information and photosMutton snapper, caught in 100 ft of water off Juno Beach, FL
* {{Taxonbar, from=Q2692345 mutton snapper Fish of the Caribbean Taxonomy articles created by Polbot mutton snapper Taxa named by Georges Cuvier