The

Brachycera The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristic ...

n infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known

flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwing ...

. It includes a number of the most familiar flies, such as the

housefly The housefly (''Musca domestica'') is a fly of the suborder Cyclorrhapha. It is believed to have evolved in the Cenozoic Era, possibly in the Middle East, and has spread all over the world as a commensal of humans. It is the most common f ...

, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal

arista Arista may refer to: Organizations *Arista Networks, a software defined networking company *Arista Records, an American record label, division of Sony Music **Arista Nashville, a record label specializing in country music *Arista (honor society) ...

. Their bodies are often highly setose, and the pattern of setae is often taxonomically important. The

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...

e of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the

pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...

e are formed inside the

exoskeleton An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton ( endoskeleton) in for example, a human. In usage, some of the ...

of the last

instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow o ...

; exit from this

puparium A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...

is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name,

Cyclorrhapha Cyclorrhapha is an unranked taxon within the infraorder Muscomorpha. They are called "Cyclorrhapha" ('circular-seamed flies') with reference to the circular aperture through which the adult escapes the puparium. This is a circumscriptional name t ...

Classification

The name

is used, in various modern classifications, to represent either a subgroup within the infraorder Muscomorpha, or simply a rankless group within the Brachycera. In either case, the Empidoidea are the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the Cyclorrhapha. In the present classification, as the term Muscomorpha is used to refer to the sister taxon of the Empidoidea, the names "Muscomorpha" and "Cyclorrhapha" are effectively synonymous (though not entirely interchangeable: for nomenclatural purposes, it is always considered better if the endings of names of similar rank are consistent, and since all the other infraordinal names end in "-morpha", the use of "Cyclorrhapha" as an infraordinal name would be inconsistent). In the

Tree of Life Web Project The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site ...

, the name "Muscomorpha" refers to the Asilomorpha plus the Cyclorrhapha. *Section Aschiza **Superfamily Platypezoidea **Superfamily

Syrphoidea The Syrphoidea are a superfamily of flies containing only two families under present classification, one of which ( Syrphidae) has a great number of the most common and familiar flies. One of these familiar flies is ''Eristalis tenax ''Eristali ...

*Section

Schizophora The Schizophora are a section of true flies containing 78 families, which are collectively referred to as muscoids, although technically the term "muscoid" should be limited to flies in the superfamily Muscoidea; this is an example of informal, ...

**Subsection

Acalyptratae The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, a ...

***Superfamily

Conopoidea The Conopidae, usually known as the thick-headed flies, are a family of flies within the Brachycera suborder of Diptera, and the sole member of the superfamily Conopoidea. Flies of the family Conopidae are distributed worldwide in all the biog ...

***Superfamily

Tephritoidea The Tephritoidea are a superfamily of flies. It has over 7,800 species, the majority of them in family Tephritidae. The following families are included: * Ctenostylidae * Eurygnathomyiidae * Lonchaeidae - lance flies * Pallopteridae — flutt ...

***Superfamily Nerioidea ***Superfamily

Diopsoidea The Diopsoidea are a small but diverse cosmopolitan superfamily of acalyptrate muscoids, especially prevalent in the tropics. Some flux exists in the family constituency of this group, with the Strongylophthalmyiidae and Tanypezidae formerly bei ...

***Superfamily Sciomyzoidea ***Superfamily

Sphaeroceroidea Sphaeroceroidea is a superfamily of flies. It includes the cosmopolitan families of Sphaeroceridae (small dung flies), Heleomyzidae, and Chyromyidae, as well as a few smaller groups. It has about 2,600 species. Description Sphaeroceroids tend ...

***Superfamily

Lauxanioidea The Lauxanioidea are a superfamily of flies that includes the two large families, the Lauxaniidae and Chamaemyiidae, and the small family Celyphidae. Generally, they are small to medium, densely populated, coloured flies. The Chamaemyiidae liv ...

***Superfamily Opomyzoidea ***Superfamily Ephydroidea ***Superfamily Carnoidea **Subsection Calyptratae ***Superfamily Muscoidea ***Superfamily Oestroidea ***Superfamily Hippoboscoidea

External links

The Tree of Life Web Project: Brachycera
{{Taxonbar, from=Q240514 Insect infraorders