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Glover's pika (''Ochotona gloveri'') is a species of
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
in the family
Ochotonidae A pika ( , or ) is a small, mountain-dwelling mammal native to Asia and North America. With short limbs, a very round body, an even coat of fur, and no external tail, they resemble their close relative the rabbit, but with short, rounded ears. ...
. It was first described in 1922, by Michael Rogers Oldfield Thomas. The summer dorsal
pelage A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
is grayish rufous, grayish brown, or tea brown in colour. The winter pelage is similar to the summer pelage, but is lighter in tone.
Endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, it is found in high altitudes of northeastern
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, southwestern
Qinghai Qinghai is an inland Provinces of China, province in Northwestern China. It is the largest provinces of China, province of China (excluding autonomous regions) by area and has the third smallest population. Its capital and largest city is Xin ...
, western
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
, and northwestern
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
. It is a generalist
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
, and is known to construct haypiles. It is rated as a species of 
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
 by the 
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
. It is also regionally red listed as least concern in China.


Taxonomy

Glover's pika was first described in 1922 by the British
zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
Michael Rogers Oldfield Thomas, in the 
Annals and Magazine of Natural History The ''Journal of Natural History'' is a scientific journal published by Taylor & Francis focusing on entomology and zoology. The journal was established in 1841 under the name ''Annals and Magazine of Natural History'' (''Ann. Mag. Nat. Hist.'') ...
. It belongs to the subgenus ''Conothoa''. There are four recognized subspecies: * ''O. g. brookei'' * ''O. g. calloceps'' * ''O. g. gloveri'' * '' O. g. muliensis'' Its taxonomic placement has been controversial and subject to change. In 1951, Ellerman and Terence Morrison-Scott assigned ''O. g. brookei'' as a subspecies of the Turkestan red pika ('' Ochotona rutila''), but it is now treated as a subspecies of Glover's pika. Pen et al., in 1962, described the Muli pika ('' Ochotona muliensis'') as a new form of the Glover's pika; the Muli pika was afterwards considered an independent species based on skull morphology, but it is currently considered a subspecies of the Glover's pika. The Glover's pika was earlier treated as a subspecies of the Turkestan red pika, but it is now considered as an independent species based on the skull morphology,
pelage A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
color, and
zoogeography Zoogeography is the branch of the science of biogeography that is concerned with geographic distribution (present and past) of animal species. As a multifaceted field of study, zoogeography incorporates methods of molecular biology, genetics, mo ...
. It had also been treated as a subspecies of the Chinese red pika ('' Ochotona erythrotis'') partly because of its assignment under the Turkestan red pika. According to morphological and molecular studies, the Glover's pika is closely related to the Chinese red pika and the two are
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
s. In 1948, Argyropulo described the species ''Ochotona kamensis'', and explained it to be the larger and more differentiated than the Chinese red pika and the Glover's pika. In 1985, after observing striking similarities between their skull structures and coat colors, Feng and Zheng synonymized ''Ochotona kamensis'' with ''Ochotona gloveri''. In 1988, however, Erbajeva synonymized ''Ochotona kamensis'' with ''Ochotona gloveri brookei'', and retained the name kamensis, as she did not use the
principle of priority Priority is a principle in Taxonomy (biology), biological taxonomy by which a valid scientific name is established based on the oldest available name. It is a decisive rule in Botanical nomenclature, botanical and zoological nomenclature to recogn ...
which recognizes the first valid application of a name to a plant or animal.


Description

The Glover's pika measures in length, and weighs . It has a long skull with its dorsal side being arch shaped due to the oblique backward sloping of the
parietal bone The parietal bones ( ) are two bones in the skull which, when joined at a fibrous joint known as a cranial suture, form the sides and roof of the neurocranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four bord ...
. The frontal bone protrudes forward slightly, and has two oval alveoli above it. The summer dorsal pelage is grayish rufous, grayish brown, or tea brown in colour. The head is orange or pale brown from the
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...
to the frontal bone. The ears are large, measuring in length, are thin haired, and light chestnut, orange, or orange brown colored. The winter pelage is similar to the summer pelage, but is lighter in tone. The ventral and upper portion of the feet are dull gray or grayish white. The hindfeet are long. The anterior palatal foramen and the palatal foramen are detached or adjoined. The
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
''O. g. gloveri'' has a dark gray cheek and smoky yellow rostrum. ''O. g. brookei'' has an orange russet tipped nose, and light gray spotted above the eyes. The Glover's pika is similar to the Chinese red pika, but it is distinguished by its smaller auditory bullae, more slender rostrum, and longer nasals than the Chinese red pika. It also does not have the reddish mantle as possessed by the Chinese red pika.


Distribution and habitat

The Glover's pika species is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. It is found in high altitudes of northeastern
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, southwestern
Qinghai Qinghai is an inland Provinces of China, province in Northwestern China. It is the largest provinces of China, province of China (excluding autonomous regions) by area and has the third smallest population. Its capital and largest city is Xin ...
, western
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
, and northwestern
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
. The nominate subspecies ''O. g. gloveri'' is found in western Sichuan, ''O. g. brookei'' is found in southwestern Qinghai and northeastern Tibet, and ''O. g. calloceps'' is found in northwestern Yunnan and eastern Tibet. It is a rock-dwelling pika which usually inhabits
scree Scree is a collection of broken rock fragments at the base of a cliff or other steep rocky mass that has accumulated through periodic rockfall. Landforms associated with these materials are often called talus deposits. The term ''scree'' is ap ...
deposits and rocky clefts which is bordered by woodlands in most of its range. It is also known to live in adobe walls of lamaseries and villages, and does not live in grazing or agricultural habitats. It is typically found in high elevations ranging from from sea level but, in Sichuan, it may occur at lower altitudes of .


Behavior and ecology

Very little is known about the Glover's pika's behavior, ecology, and reproduction. It is a generalist
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
, and is known to construct haypiles.


Status and conservation

Since 1996, the Glover's pika is rated as a species of 
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
 on the  IUCN Red List of Endangered Species. This is because it is a widespread species, and it occurs in remote areas of China which are unlikely to be affected by human activities. The current state of its population trend is unclear, as no natural historic or ecological investigation has been done on this species. However, being locally poisoned and trapped is a major threat to the Glover's pika populations. It is also regionally red listed as a species of least concern in China.


References


Further reading

*


External links

* {{Taxonbar, from=Q1761829 Pikas Mammals of China Endemic fauna of China Mammals described in 1922 Taxa named by Oldfield Thomas Taxonomy articles created by Polbot