Mud Snake
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The mud snake (''Farancia abacura'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of non
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
, semiaquatic,
colubrid Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. Colu ...
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the southeastern United States.


Description

The mud snake usually grows to a total length (including tail) of 40 to 54 inches (1-1.4 m),Missouri Department of Conservation (2013)
Western Mud Snake
MDC Online. Accessed May 18, 2013
with the record total length being slightly over 80 inches (2 m).The University of Georgia (2008)
Mud Snake
The University of Georgia: Museum of Natural History. Accessed April 23, 2011.
This species is sexually dimorphic in size. Female adults are larger than males in total length. The upperside of the mud snake is glossy black. The underside is red and black, and the red extends up the sides to form bars of reddish-pink. Although, some have a completely black body with slightly lighter black spots instead of the common reddish colors. The heavy body is cylindrical in cross section, and the short tail has a terminal spine. The head scalation is distinctive in that there is only one internasal scale, no
preocular In scaled reptiles, the ocular scales are those forming the margin of the eye.Wright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . The name originates from the term which is Latin f ...
scale, and one anterior temporal scale. The
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are smooth, and are arranged in 19 rows at midbody. There are 168–208
ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
and 31–55
subcaudal scales In snakes, the subcaudal scales are the enlarged plates on the underside of the tail. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates, a Division of Cornell University Press. (7t ...
. The anal plate is divided. ''Farancia abacura'' is the only species that belongs to the ''Farancia'' genus, with two different subspecies within itself.


Subspecies

There are two recognized
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of ''Farancia abacura'', including the
nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
: *''
Farancia abacura abacura ''Farancia'' is a genus of colubrid snakes. It consists of two species, one commonly referred to as the rainbow snake (''F. erytrogramma'') and the other commonly referred to as the mud snake (''F. abacura''). Both species are native to the sout ...
'' ( Holbrook, 1836) – eastern mud snake *''
Farancia abacura reinwardtii The mud snake (''Farancia abacura'') is a species of nonvenomous, semiaquatic, colubrid snake endemic to the southeastern United States. Description The mud snake usually grows to a total length (including tail) of 40 to 54 inches (1-1.4 m),Misso ...
'' ( Schlegel, 1837) – western mud snake


Geographic range

The mud snake is found in the southeastern United States, in the states of Alabama, Arkansas, Florida, Georgia, Illinois, Kentucky, Louisiana, Mississippi, Missouri, North Carolina, Oklahoma, South Carolina, Tennessee, Texas, and Virginia.Willson JD (2006)
Mud Snake (''Farancia abacura'')
Savannah River Ecology Laboratory. Accessed April 23, 2011.


Habitat

''F. abacura'' inhabits the edges of
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a strea ...
s and cypress swamps, among dense vegetation or under ground debris, using wet conditions to burrow itself into the mud. It is almost fully aquatic and rarely leaves the water, except to lay eggs, hibernate, or during drought to escape drying wetlands. After heavy rain fall, its home range may increase to take advantage of new food sources. It occupies aquatic habitats with freshwater or brackish waters. For hibernation, they commonly use cavities in soil or old tree stumps.


Behavior

The mud snake is mostly
aquatic Aquatic means relating to water; living in or near water or taking place in water; does not include groundwater, as "aquatic" implies an environment where plants and animals live. Aquatic(s) may also refer to: * Aquatic animal, either vertebrate ...
and
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
. It preys mostly on giant aquatic salamanders in the genera '' Siren'' and ''
Amphiuma ''Amphiuma'' is a genus of aquatic salamanders from the United States, the only Extant taxon, extant genus within the family (biology), family Amphiumidae . They are colloquially known as amphiumas. They are also known to fishermen as "conger eel ...
'', but it also eats other amphibians. They are known to use their sharply pointed tails to prod
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
items, leading to the nickname "stinging snake", although their tail is not a
stinger A stinger (or sting) is a sharp organ found in various animals (typically insects and other arthropods) capable of injecting venom, usually by piercing the epidermis of another animal. An insect sting is complicated by its introduction of ve ...
and cannot sting. Enlarged teeth occur at the rear of the upper jaw, which presumably help to hold slippery prey. Upon being disturbed, mud snakes will sometimes tuck their head beneath their coils and expose the red underside on the tail as a warning display.


Reproduction

Breeding of ''F. abacura'' takes place in the spring, mostly in the months of April and May. During copulation, the female will wind itself around the male. They can stay in this position for over a day. Eight weeks after mating, the female lays 4 to 111
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
in a nest dug out of moist soil, sometimes in alligator nests. The nests of this species can commonly be found in the ground under debris, but it is not limited to those areas. These nests are cavities in sandy soil with the eggs laid within. Some eggs can even be embedded into the wall. There is a positive correlation between body length and clutch size, with larger females having larger clutch sizes. She will remain with her eggs until they hatch, in the fall, usually September or October. Although unhatched eggs have not been found in the winter or spring, many juvenile mudsnakes are captured entering wetlands in the spring, most likely from clutches deposited and hatched in the preceding late summer or autumn. It is thought that mudsnake hatchlings either enter aquatic habitats in the autumn or delay entering them until the spring, but it is not known if they remain in a terrestrial nest or disperse into terrestrial habitats during this time.


Cultural significance

The mud snake is one of a few animals which may be the origin of the hoop snake myth. J.D. Willson writes: The hoop snake myth has also been attributed to the coachwhip snake.


References


External links

*
Illinois Natural History Survey: ''Farancia abacura''"Black Snakes": Identification and Ecology
– University of Florida fact sheet


Further reading

* Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp., 657 color plates. . (''Farancia abacura'', pp. 609–610 + Plate 492). * Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition''. Boston: Houghton Mifflin. xviii + 429 pp. (hardcover), (paperback). (''Farancia abacura'', pp. 176–177 + Plate 25 + Map 138). *Conant R, Bridges W (1939). ''What Snake Is That?: A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (With 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century. Frontispiece map + viii + 163 pp. + Plates A-C, 1–32. (''Farancia abacura'', pp. 33–36 + Plate 3, Figure 7). * * Holbrook JE (1836). ''North American Herpetology; or, A Description of the Reptiles Inhabiting the United States. Vol. I.'' Philadelphia: J. Dobson. 120 pp. (''Coluber abacurus'', new species, pp. 119–120). * *Morris PA (1948). ''Boy's Book of Snakes: How to Recognize and Understand Them''. A volume of the Humanizing Science Series, edited by
Jacques Cattell Jaques (Jack) Cattell (2 June 1904 in Garrison, New York – 19 December 1961) was an American publisher and founder of a company bearing his name, "Jaques Cattell Press, Inc.," based in Lancaster, Pennsylvania. Jaques Cattell Press, Inc. The Sci ...
. New York: Ronald Press. New York. viii + 185 pp. ("The Mud Snake", ''Farancia abacura'', pp. 87–88, 179). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp., 47 plates, 207 figures. (''Farancia abacura'', pp. i, 405–406 + Plate 39). * Schlegel H (1837). ''Essai sur la physionomie des serpens.'' Amsterdam: M.H. Schonekat. Amsterdam. ''Partie Générale.'' xxviii + 251 pp. (''Homalopsis reinwardtii'', new species, p. 173). AND ''Partie Descriptive.'' 606 + xvi pp. (''Homalopsis reinwardtii'', pp. 357–358). (in French). * Schmidt KP, Davis DD (1941). ''Field Book of Snakes of the United States and Canada''. New York: G.P. Putnam's Sons. 365 pp. (''Farancia abacura'', pp. 106–108, Figure 22 + Plate 9). * Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Itahaca and London: Comstock. 1,105 pp. (in 2 volumes) (''Farancia abacura'', pp. 271–280, Figures 84–85, Map 25). * Zim HS, Smith HM (1956). ''Reptiles and Amphibians: A Guide to Familiar American Species: A Golden Nature Guide''. New York: Simon and Schuster. 160 pp. (''Farancia abacura'', pp. 75, 156). {{Taxonbar, from=Q517458 Farancia Reptiles of the United States Fauna of the Southeastern United States Reptiles described in 1836 Semiaquatic animals Taxa named by John Edwards Holbrook