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The Eurasian woodcock (''Scolopax rusticola'') is a medium-small wading bird found in temperate and subarctic
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
. It has cryptic camouflage to suit its woodland habitat, with reddish-brown upperparts and buff-coloured underparts. Its eyes are set far back on its head to give it 360-degree vision and it probes in the ground for food with its long, sensitive bill, making it vulnerable to cold weather when the ground remains frozen. The male performs a courtship flight known as "roding" at dusk in spring. It is widely believed that the female will sometimes carry chicks between her legs whilst flying, though evidence of this is purely anecdotal. The world population is estimated to be 14 million to 16 million birds.


Taxonomy

The Eurasian woodcock was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the current
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Scolopax rusticola''. The genus name is
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
for a
snipe A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/ camouflaging plumage. ''Gallinago'' snipe have a nearly ...
or woodcock. The specific epithet ''rusticola'' is the Latin name of a gamebird mentioned by
Pliny the Elder Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...
and
Marcus Valerius Martialis Marcus Valerius Martialis (known in English as Martial ; March, between 38 and 41 AD – between 102 and 104 AD) was a Roman and Celtiberian poet born in Bilbilis, Hispania (modern Spain) best known for his twelve books of ''Epigrams'', pub ...
. It was possibly a
grouse Grouse are a group of birds from the order (biology), order Galliformes, in the family (biology), family Phasianidae. Grouse are presently assigned to the Tribe (biology), tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetr ...
. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised. Its closest relatives are the other woodcocks in the genus '' Scolopax''.


Description

Adults are in length, including the long straight bill, and have a wingspan. The Eurasian woodcock has cryptic camouflage to suit its woodland habitat, with intricately patterned reddish-brown upperparts and buff underparts. The head is barred with black, not striped like that of its close relatives, the
snipe A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/ camouflaging plumage. ''Gallinago'' snipe have a nearly ...
. It has large eyes located high on the sides of its head, giving it 360-degree monocular vision. The wings are rounded and the base of the bill is flesh-coloured with a dark tip. The legs vary from grey to pinkish. The species is
sexually dimorphic Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
, with the male much larger than the female, although the sexes cannot be separated in the field.


Distribution and habitat

About one third of the world's Eurasian woodcock breed in Europe, with more than 90% of the continent's population breeding in
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
and
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...
. Their breeding range stretches from Fennoscandia to the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
and
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
and from western Europe to Russia. The population on the Azores is isolated from mainland Europe, resulting in subtle genetic differentiation. This species is the woodcock found through most of temperate and subarctic Eurasia. Northern and Asian populations
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
to southern Europe or the
Indian Subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
, respectively. Birds in milder western European countries and on Atlantic islands are resident. The populations breeding in north-west and southern Europe are mostly sedentary. The birds' spring migration commences in February; breeding territories are reached between March and May. Spring migration movements of woodcock are influenced by weather conditions, however this does not have a major effect on their subsequent breeding success. The Eurasian woodcock has a large range, with an estimated Global Extent of Occurrence of 10 million square kilometres and a population of an estimated 15 million to 16 million birds. Because of its large range, stable population trend and large population size, the species has been evaluated as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. The most important threat to the population of the Eurasian woodcock in the breeding range is the increased fragmentation of its woodland habitat. At other times of year, a reduction in the amount of permanent grassland and an increase in the intensification of farming are also threats, and the species' susceptibility to
avian influenza Avian influenza, also known as avian flu or bird flu, is a disease caused by the influenza A virus, which primarily affects birds but can sometimes affect mammals including humans. Wild aquatic birds are the primary host of the influenza A viru ...
may have an impact in future. The Eurasian woodcock's required breeding habitat is large, unfragmented areas of broadleaved deciduous or mixed broadleaved and coniferous forest, with dense undergrowth of plants such as
brambles ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
,
holly ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...
,
hazel Hazels are plants of the genus ''Corylus'' of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family, Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K ...
,
gorse ''Ulex'' (commonly known as gorse, furze, or whin) is a genus of flowering plants in the family Fabaceae. The genus comprises about 20 species of thorny evergreen shrubs in the subfamily Faboideae of the pea family Fabaceae. The species are na ...
,
bracken Bracken (''Pteridium'') is a genus of large, coarse ferns in the family (biology), family Dennstaedtiaceae. Ferns (Pteridophyta) are vascular plants that undergo alternation of generations, having both large plants that produce spores and small ...
, or
bilberry Bilberries () are Eurasian low-growing shrubs in the genus ''Vaccinium'' in the flowering plant family Ericaceae that bear edible, dark blue berries. They resemble but are distinct from North American blueberries. The species most often referre ...
. On a small scale, Eurasian woodcock appear to prefer woods containing birch and were less likely to be found in beech dominated woods. They also prefer woods further away from urban areas. Breeding territories must include a mix of dry, warm resting places, damp areas for feeding, and clearings for flight. In larger woods, wide "rides" (open tracks through the wood) and small clearings are important. In winter, Eurasian woodcock also use
scrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
during the day but in freezing weather they may use intertidal mud.


Behaviour

Eurasian woodcock are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal (active during dawn), vespertine (biology), vespertine/vespertinal (active during dusk), or both. This is distinguished from diurnalit ...
(most active at dawn and dusk) and rarely active during the day unless flushed, when they fly off with a whirring wing noise. The flight is somewhat owl- or bat-like; woodcock fly fast and directly while migrating or crossing open country, but fly erratically with twisting and fluttering once in woodland. They are usually solitary and migrate singly, but may congregate when weather or geographical conditions force them to do so. This species is parasitised by the
moorhen flea The moorhen flea (''Dasypsyllus gallinulae'') is a flea originating from South America. It is now globally widespread. It is a large flea, easily identified because the male has two heavy horn-like spines on one of the genital flaps, and the fema ...
, ''Dasypsyllus gallinulae''.


Breeding

The male performs a courtship display flight called 'roding' at dusk between April and June. A
mating call A mating call is the auditory signal used by animals to attract mates. It can occur in males or females, but literature is abundantly favored toward researching mating calls in females. In addition, mating calls are often the subject of mate choi ...
is performed during courtship display flights: ''orr, orr, orr, pist''. A study of
spectrogram A spectrogram is a visual representation of the spectrum of frequencies of a signal as it varies with time. When applied to an audio signal, spectrograms are sometimes called sonographs, voiceprints, or voicegrams. When the data are represen ...
s of the vocalisations of male Eurasian woodcock showed that 95% of calls could be attributed correctly to individual birds, and concluded that counting roding males was a suitable method of monitoring the population. Roding surveys are now used for population estimates in a range of countries including Britain, Switzerland, France and Russia. Eurasian woodcock nest on the ground in low cover in woodland or tall heather. The nest is a lined cup or a slight hollow lined with dead leaves and other plant material. A single brood of one or two white or creamy eggs with light brown and grey blotches is laid; once the clutch is complete, incubation is performed by the female for between 21 and 24 days. The eggs measure and weigh , of which 5% is shell. The downy,
precocial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
young leave the nest immediately and fledge after 15–20 days, although they can fly short distances after 10 days. When threatened, the mother bird can fly whilst carrying small chicks between her legs, body and tail, in her claws or on her back. This behaviour is rarely witnessed.


Food and feeding

Eurasian woodcock forage in soft soil in thickets, usually well hidden from sight. They mainly eat earthworms, but also insects and their larvae, freshwater molluscs and some plant seeds. Because they rely on probing into the ground to find food, they are vulnerable to cold winter weather when the ground remains frozen. During the cold British winter of 1962–63, starving woodcock were found feeding in urban areas and some were even forced to eat bird seed.
Nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
feeding occurs in frost-free
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
or long-rotation grass
meadow A meadow ( ) is an open habitat or field, vegetated by grasses, herbs, and other non- woody plants. Trees or shrubs may sparsely populate meadows, as long as they maintain an open character. Meadows can occur naturally under favourable con ...
s; fields with woodland within are preferred to allow roosting during the day. There are only low densities of Eurasian woodcock in predominantly arable areas. In a three-year scientific study based in
Brittany Brittany ( ) is a peninsula, historical country and cultural area in the north-west of modern France, covering the western part of what was known as Armorica in Roman Gaul. It became an Kingdom of Brittany, independent kingdom and then a Duch ...
,
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
, 65 radio-tagged woodcock and their habitat preferences were monitored. During the day, the birds chose areas of forest with humus which held large numbers of earthworms and with a dense scrub layer to provide protection, and hedges with scrub and trees were also found to be important. At night, the woodcock selected grazed pastures which had five times the
biomass Biomass is a term used in several contexts: in the context of ecology it means living organisms, and in the context of bioenergy it means matter from recently living (but now dead) organisms. In the latter context, there are variations in how ...
of earthworms compared to tilled land.


Relationship to humans


Hunting

In many countries woodcock are hunted as game, and their size, speed, and
flight pattern ''Flight Pattern'' is a contemporary ballet choreographed by Crystal Pite, set to the first movement of Henryk Górecki's Symphony No. 3. It premiered at the Royal Opera House, London, on 16 March 2017, making Pite the first woman to cho ...
makes them a very challenging shot. Anyone shooting a witnessed "right and left"—in which two woodcock are killed with consecutive shots from a shotgun, without the gun being lowered or reloaded—can be admitted to the ''
Shooting Times ''Shooting Times & Country Magazine'', more commonly known as the ''Shooting Times'', is a Great Britain, British shooting, fieldsports, and conservation magazine, published by Fieldsports Press Ltd. The magazine also features articles on fishing ...
'' Woodcock Club. In 1821
William Bingley William Bingley (January 1774 – 11 March 1823) was an English cleric, naturalist and writer. Life Bingley was born at Doncaster, and left an orphan at an early age. In 1795 he entered Peterhouse, Cambridge, and took the degree of B.A. in 1799, a ...
described a method of trapping Eurasian woodcock formerly used in northern England: "Long parallel rows of stones or stick, four or five inches high, were made in moonlight nights on the commons frequented by them. In these rows several intervals or gateways were left in which the traps were placed. When the birds, running about in search of food, came to one of these rows, they did not usually cross it, but ran along the side till they arrived at the gateways, which they entered, and in which they were caught." The effect of hunting on the species' European breeding population is "little understood". A study conducted in
Brittany Brittany ( ) is a peninsula, historical country and cultural area in the north-west of modern France, covering the western part of what was known as Armorica in Roman Gaul. It became an Kingdom of Brittany, independent kingdom and then a Duch ...
, France, explored the factors affecting the populations of wintering Eurasian woodcock. The mortality and survival rates of 98 radio-tagged birds were monitored in a hunted area and in an adjoining hunting-free reserve. The scientists found rates of survival of 0.86 for adults and 0.67 for juveniles, with the probability of survival increasing as birds spent more time in the reserve.
Matrix population models Matrix population models are a specific type of population model that uses matrix algebra. Population models are used in population ecology to model the dynamics of wildlife or human populations. Matrix algebra, in turn, is simply a form of algebr ...
were used and the conclusion was that "such low survival probabilities cannot sustain long-term viable populations. These results call for caution in the harvesting of woodcock populations wintering in western France and could be a forewarning of a decline." In the United Kingdom, surveys are being conducted to improve knowledge of the number of breeding Eurasian woodcock and the structure of their habitat.
Isotope Isotopes are distinct nuclear species (or ''nuclides'') of the same chemical element. They have the same atomic number (number of protons in their Atomic nucleus, nuclei) and position in the periodic table (and hence belong to the same chemica ...
analysis of feathers from Eurasian woodcock shot in the UK is also being performed to establish the origins of the birds.


Gastronomy

Eurasian woodcock are considered at their best for culinary purposes from October to December. After shooting, the birds are hung for 4–8 days to tenderise; the meat can be eaten as a starter, savoury, or for breakfast. A bird serves one person. They are not gutted before cooking; the head, skinned with eyes removed, is left on and the beak used to truss the bird. A recipe for roast woodcock was featured in ''
Mrs Beeton's Book of Household Management ''Mrs. Beeton's Book of Household Management'', also published as ''Mrs. Beeton's Cookery Book'', is an extensive guide to running a household in Victorian Britain, edited by Isabella Beeton and first published as a book in 1861. Previously ...
'', but despite its name, the
Victorian Victorian or Victorians may refer to: 19th century * Victorian era, British history during Queen Victoria's 19th-century reign ** Victorian architecture ** Victorian house ** Victorian decorative arts ** Victorian fashion ** Victorian literatur ...
dish named
Scotch woodcock Scotch woodcock is a British savoury dish consisting of creamy, lightly-scrambled eggs served on toast that has been spread with anchovy paste or Gentleman's Relish, and sometimes topped with chopped herbs and black pepper. Scotch woodcock was s ...
consists of
scrambled eggs Scrambled eggs is a dish made from eggs (usually chicken eggs), where the whites and yolks have been stirred, whipped, or beaten together (typically with salt, butter or oil, and sometimes water or milk, or other ingredients), then heated so ...
on toast with
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
. Bingley wrote that (in the United Kingdom) "Few birds are so much in esteem for the table as these", although he also reported that woodcock meat was not eaten in Sweden or Norway as it was thought unwholesome, but that the eggs were taken and sold in "great numbers" in
Stockholm Stockholm (; ) is the Capital city, capital and List of urban areas in Sweden by population, most populous city of Sweden, as well as the List of urban areas in the Nordic countries, largest urban area in the Nordic countries. Approximately ...
and
Gothenburg Gothenburg ( ; ) is the List of urban areas in Sweden by population, second-largest city in Sweden, after the capital Stockholm, and the fifth-largest in the Nordic countries. Situated by the Kattegat on the west coast of Sweden, it is the gub ...
. In
Nicolas Freeling Nicolas Freeling (born Nicolas Davidson; 3 March 1927 – 20 July 2003), was a British crime novelist, best known as the author of the "Van der Valk" series of detective novels. A television series based on the character, ''Van der Valk'', was pro ...
's ''Kitchen Book'' he narrates the lavish dinner described in
Kipling Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)''The Times'', (London) 18 January 1936, p. 12. was an English journalist, novelist, poet, and short-story writer. He was born in British India, which inspired much of his work. ...
's '' Sea Constables'', which features woodcock as a savoury, and explains that 'the woodcock has no gall, and is not cleaned ..the creamy inside parts are spread on toast ..no Edwardian gentleman would have resisted this for one minute.'


Folklore and culture

In the United Kingdom, the early arrival of migrant woodcock in autumn was said to mean a good harvest, especially if they stayed until spring. It used to be thought that Eurasian woodcock flew to the moon during the months when they were not seen and the first full moon in November, when large numbers arrive on the British coast, is sometimes described as the 'woodcock moon'. Reaney and Wilson wrote that the English surname "Woodcock" developed as a nickname from the bird, meaning 'a fool, simpleton or dupe'. In
William Shakespeare William Shakespeare ( 23 April 1564 – 23 April 1616) was an English playwright, poet and actor. He is widely regarded as the greatest writer in the English language and the world's pre-eminent dramatist. He is often called England's nation ...
's play ''
Love's Labours Lost ''Love's Labour's Lost'' is one of William Shakespeare's early comedies, believed to have been written in the mid-1590s for a performance at the Inns of Court before Queen Elizabeth I. It follows the King of Navarre and his three companions as ...
'', Berowne describes himself and his friends as 'four woodcocks in a dish', after discovering they have all fallen in love when they have sworn not to. An old folk name for the
goldcrest The goldcrest (''Regulus regulus'') is a very small passerine bird in the kinglet family. Its colourful golden Crest (feathers), crest feathers, as well as being called the "king of the birds" in European folklore, gives rise to its Englis ...
was the "woodcock pilot" because of the mistaken belief that the warblers rode in the feathers of migrating woodcock. In ''
Hamlet ''The Tragedy of Hamlet, Prince of Denmark'', often shortened to ''Hamlet'' (), is a Shakespearean tragedy, tragedy written by William Shakespeare sometime between 1599 and 1601. It is Shakespeare's longest play. Set in Denmark, the play (the ...
'', Polonius refers to Hamlet's vows as "springes to catch woodcocks". The 'pin feather' of the Eurasian woodcock (the
covert feather A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
of the outermost
primary Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Work ...
) can be used as a fine paintbrush for miniatures, or as a delicate tool for moving tiny objects (e.g. specks of dust). As a painting tool, the feather has been described as "far from ideal, holding very little paint, resisting water and wearing down quickly at the tip." The Eurasian woodcock has been the subject of
postage stamp A postage stamp is a small piece of paper issued by a post office, postal administration, or other authorized vendors to customers who pay postage (the cost involved in moving, insuring, or registering mail). Then the stamp is affixed to the f ...
s from several countries. A woodcock serves as a symbolic element in the novel '' The Birds'' by writer
Tarjei Vesaas Tarjei Vesaas (20 August 1897 – 15 March 1970) was a Norwegian poet and novelist. Vesaas is widely considered to be one of Norway's greatest writers of the twentieth century and perhaps its most important since World War II. Vesaas' work is c ...
.


References


External links


Eurasian woodcock photos
a
Oiseaux.net
*
Woodcock
a
RSPB Birds by Name

Ageing and sexing (PDF; 2.9 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze


a
Ornithos.de
* * * * * {{Authority control
Eurasian woodcock The Eurasian woodcock (''Scolopax rusticola'') is a medium-small wader, wading bird found in temperate and subarctic Palearctic realm, Eurasia. It has Camouflage#Cryptic coloration in nature, cryptic camouflage to suit its woodland habitat, with ...
Birds of Eurasia Wading birds Birds described in 1758 Game birds Taxa named by Carl Linnaeus Birds of Iran