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The Cape gannet (''Morus capensis'') is a large
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
of the
gannet Gannets are seabirds comprising the genus ''Morus'' in the family Sulidae, closely related to boobies. They are known as 'solan' or 'solan goose' in Scotland. A common misconception is that the Scottish name is 'guga' but this is the Gaelic n ...
family,
Sulidae The bird family Sulidae comprises the gannets and boobies. Collectively called sulids, they are medium-large coastal seabirds that plunge-dive for fish and similar prey. The 10 species in this family are often considered congeneric in older so ...
. They are easily identified by their large size, black and white
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
and distinctive yellow crown and hindneck. The pale blue bill is pointed with fine serrations near the tip; perhaps because of the depth and speed of the gannet's dive when fishing (depending on altitude, gannets hit the water at speeds of between ), its beak has no external nostrils into which the water might be forced. The widely mentioned maximum speed of 120 km/h has never been measured and published.


Taxonomy

German naturalist
Hinrich Lichtenstein Martin H nrich Carl Lichtenstein (10 January 1780 – 2 September 1857) was a German physician, List of explorers, explorer, botanist and zoologist. He explored parts of southern Africa and collected natural history specimens extensively and ...
described the Cape gannet in 1823. The
Sulidae The bird family Sulidae comprises the gannets and boobies. Collectively called sulids, they are medium-large coastal seabirds that plunge-dive for fish and similar prey. The 10 species in this family are often considered congeneric in older so ...
, the
gannet Gannets are seabirds comprising the genus ''Morus'' in the family Sulidae, closely related to boobies. They are known as 'solan' or 'solan goose' in Scotland. A common misconception is that the Scottish name is 'guga' but this is the Gaelic n ...
s and
boobies A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fre ...
, appeared about 30 million years ago. Early Sulidae fossils most resembled the boobies, although they were more aquatic, with the gannets splitting off later, about 16 million years ago. The gannets evolved in the northern hemisphere, later colonising the southern oceans. The most ancient extant species may be the
Abbott's booby Abbott's booby (''Papasula abbotti'') is an endangered seabird of the Sulidae, sulid family, which includes gannets and Booby, boobies. It is a large booby and is placed within its own monotypic genus. It was first identified from a specimen col ...
, possibly the sole survivor of an otherwise extinct separate lineage. A 2011 genetic study of
nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics * Nuclear space *Nuclear ...
and
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
suggests that the ancestor of the gannets arose around 2.5 million years ago before splitting into northern and southern lineages. The latter then splitting into the Cape and
Australasian gannet The Australasian gannet (''Morus serrator''), also known as the Australian gannet or tākapu, is a large seabird of the booby and gannet family, Sulidae. Adults are mostly white, with black flight feathers at the wingtips and lining the trailing ...
s around 0.5 million years ago. The three gannets are generally considered to be separate species forming a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
, though they have also formerly been classified as
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
(''Sula bassanus'').


Description

When seen in flight the snow-white body with the black tail, primaries and secondaries, and dark bill makes them easy to identify. At closer range the distinctive golden crown and nape, which gradually becomes white on the neck, is noticeable. Juveniles and immatures are dark brown with a pale bill, and can resemble the dark-coloured
boobies A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fre ...
at first glance. Adults are about long and have a wingspan and weigh ''ca.'' . The Cape gannet is identified from the
Australasian gannet The Australasian gannet (''Morus serrator''), also known as the Australian gannet or tākapu, is a large seabird of the booby and gannet family, Sulidae. Adults are mostly white, with black flight feathers at the wingtips and lining the trailing ...
by the all-black tail, a longer black stripe down the throat (from underneath the beak) and more extensive black on the face, but all of these features are difficult to identify unless at close proximity. The Cape gannet is also very similar to the
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
, but the latter differs from its entirely white tail and its wings, which are only black at the tip. It also is similar to the
blue-footed booby The blue-footed booby (''Sula nebouxii'') is a marine bird native to subtropical and tropical regions of the eastern Pacific Ocean. It is one of six species of the genus ''booby, Sula'' – known as boobies. It is easily recognizable by its dis ...
(''Sula nebouxii'').


Breeding

Gannet pairs may remain together over several seasons. They perform elaborate greeting rituals at the nest, stretching their bills and necks skywards and gently tapping bills together. Cape gannets begin breeding in August or September. Typically the clutch is a single bluish
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
, which soon becomes soiled. Both parents are actively involved in the incubation process which lasts for 42 to 46 days until hatching. Gannets use their foot webs to incubate the egg. The foot webs, which are richly irrigated with blood vessels are wrapped around the egg. The hatchling is black, naked and blind, it weighs only about , but within three weeks its body mass is one third of that of an adult. At eight weeks the chick outweighs the adult, and this remains so until it becomes a fledgling at 95–105 days of age.


Feeding

Cape gannets are powerful fliers, using mainly a flap-gliding technique, which is more energy consuming than the dynamic-soaring favoured by
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...
es. As all Sulids, they are fish-eating birds that plunge-dive from considerable height.


Status and distribution


Breeding and non-breeding range

The breeding
range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...
of the Cape gannet is restricted to
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
in three islands off
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
and three islands off
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. They normally nest in large and dense colonies on flat islands or on flat ledges of the steeply sloping Mercury Island off
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
. The world population was estimated in 1996 to number about 340,000 birds, with 12% in
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
and 88% in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. The largest colony of this bird, with over 140,000 birds, is found on
Malgas Island Malgas Island is a small, , uninhabited island lying in the northern part of the entrance to Saldanha Bay, in the Western Cape province of South Africa. It lies about from the mainland in the Benguela upwelling system. It is circular in shape ...
, South Africa. Several birds have occasionally been found breeding on offshore
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
n islands, together with
Australasian gannet The Australasian gannet (''Morus serrator''), also known as the Australian gannet or tākapu, is a large seabird of the booby and gannet family, Sulidae. Adults are mostly white, with black flight feathers at the wingtips and lining the trailing ...
s, although the Cape species is never represented by more than a few pairs. The non-breeding range of the Cape gannet extends from the coastal waters off the
Gulf of Guinea The Gulf of Guinea (French language, French: ''Golfe de Guinée''; Spanish language, Spanish: ''Golfo de Guinea''; Portuguese language, Portuguese: ''Golfo da Guiné'') is the northeasternmost part of the tropical Atlantic Ocean from Cape Lopez i ...
on the west coast of Africa, to
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
on the east coast. They seldom occur farther offshore than 100 km, though records of birds more than 200 km offshore exist for both the
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
and Indian Oceans.


Population trends

Numbers of Cape gannets at the Namibian islands have declined considerably between 1956 and 2000 from 114,600 to 18,200 breeding pairs respectively, an 84% decrease in less than fifty years. This contrasts with the trends at the South African islands where numbers have increased about 4.3 times during the same period, from 34,400 to 148,000 breeding pairs.


Vagrancy

Vagrancy Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, waste picker, scavenging, or petty theft. In Western ...
to the northern
Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
has been reported for the Cape gannet on several occasions, but until 2016 was believed to be unproven. Crochet and Haas examined the status of the species in the western
Palearctic realm The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. The ...
. They list a number of historical claims which had not been accepted by national bird records committees, and analyse the single record which had hitherto been widely accepted, a juvenile recovered offshore from
Western Sahara Western Sahara is a territorial dispute, disputed territory in Maghreb, North-western Africa. It has a surface area of . Approximately 30% of the territory () is controlled by the Sahrawi Arab Democratic Republic (SADR); the remaining 70% is ...
in 1966. In relation to the latter bird, they conclude that a mistake had occurred; the original data placed the bird inland, in
Chad Chad, officially the Republic of Chad, is a landlocked country at the crossroads of North Africa, North and Central Africa. It is bordered by Libya to Chad–Libya border, the north, Sudan to Chad–Sudan border, the east, the Central Afric ...
, with the offshore Western Saharan location being the result of an attempt to correct this to a more plausible location. A retransposition of the original latitude and longitude gives a location off the
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
n coast, well within the regular wintering range of the species. As a result, they recommended that the Cape gannet be deleted from the Western Palearctic list. However, on 14 April 2016, a Cape gannet was photographed at sea off
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Administratively, it forms the largest island in the East Nusa Tenggara Province. The area is 14,250 km2. Including Komodo and Rinca islands ...
in the Azores. The species has been discussed as a possible vagrant to Britain and Ireland, and there have been a number of claimed sightings, including one in December 2020 off the
Isle of Wight The Isle of Wight (Help:IPA/English, /waɪt/ Help:Pronunciation respelling key, ''WYTE'') is an island off the south coast of England which, together with its surrounding uninhabited islets and Skerry, skerries, is also a ceremonial county. T ...
, but there is yet to be an accepted record in the Western Palearctic apart from the Azores record. One Spanish record from 1985 has been accepted as either a Cape gannet or an Australasian gannet. Individuals have also been photographed in other locations far from the species' usual range, including off Peru in 1999 and Oman in 2004.


References


Further reading

*Crawford (1997). Cape Gannet. In: ''The Atlas of Southern African Birds'' Vol. 1: Non-passerines. Harrison, J. A. Allan, D. G., Underhill. L. G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds), pp. 28–29. BirdLife South Africa, Johannesburg. *du Toit, M. & Cooper, J. (2002). Cape Gannet. In: ''Proposal for inclusion of species in Annex 2 of the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (
AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA), is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
, an agreement under UNEP/CMS)''. Avian Demography Unit, Cape Town.


External links


Species factsheet
– BirdLife International
Species text
– '' The Atlas of Southern African Birds''
Fact sheet
– Avian Demography Unit, University of Cape Town
Fact file with images and movies
– ARKive
"Beak protects during dives"
– Ask Nature
"Spinning makes safe dive"
– Ask Nature {{Taxonbar, from=Q923985 Cape gannet Birds of Southern Africa Birds of Sub-Saharan Africa Cape gannet