''Morrosaurus'' is an extinct
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of herbivorous
ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous worl ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
member of the
Elasmaria
Elasmaria is a clade of ornithopods known from Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At ...
, that lived in the late
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
in the
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
. The only known species is the
type
Type may refer to:
Science and technology Computing
* Typing, producing text via a keyboard, typewriter, etc.
* Data type, collection of values used for computations.
* File type
* TYPE (DOS command), a command to display contents of a file.
* Ty ...
''Morrosaurus antarcticus''.
Discovery
In 2002 the Argentine paleontologist
Fernando Novas reported the discovery of a partial skeleton of a euornithopod in Antarctica.
[Novas, F.E., A.V. Cambiaso, J. Lirio, & H. Núñez, 2002, "Paleobiogeografía de los dinosaurios cretácicos polares de Gondwana", ''Ameghiniana'' (Resúmenes) 39(4): 15R] In 2016 these remains were the basis for naming the type species ''Morrosaurus antarcticus'', named and described by Sebastian Rozadilla, Federico Lisandro Agnolin, Fernando Emilio Novas, Alexis Rolando Aranciaga Mauro, Matthew J. Motta, Juan Manuel Lirio Marcelo, and Pablo Isasi. The genus name refers to the site of El Morro on
James Ross Island
James Ross Island is a large island off the southeast side and near the northeastern extremity of the Antarctic Peninsula, from which it is separated by Prince Gustav Channel. Rising to , it is irregularly shaped and extends in a north–south ...
, where the remains of the species were found. The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
refers to Antarctica.
[
The holotype specimen MACN Pv 197, was found in a layer of the ]Snow Hill Island Formation
The Snow Hill Island Formation is an Early Maastrichtian geologic formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a paravian theropod ''Imperobator antarcticus''Ely & Case, 2019 have been recovered from it, a ...
(Cape Lambe, previously assigned to the Lopez de Bertodano Formation), dating to the Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
age. The remains consists in a right hind leg, including the top of the femur
The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
, the lower end of the femur, the upper part of the tibia
The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
, the underside of the tibia, the upper half of the foot, the bottom of the midfoot and the top of the first joint of the third toe.[
]
Description
The type specimen corresponds to a medium-sized animal. The descriptors were able to establish some distinctive features. Two of these are autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
, i.e. derived features that are unique. In bottom view, the greater trochanter of the femur has an undulating profile with a thick edge output and a main thin edge. The fourth metatarsal
The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...
bone has a triangular profile with a rearward projection that wraps around it and the third metatarsal.[
In addition, there is a unique combination of two features that by themselves are not unique features. In the femur, the lesser trochanter is inclined diagonally, right next to the greater trochanter. In the tibia, medial ]malleolus
A malleolus is the bony prominence on each side of the human ankle.
Each leg is supported by two bones, the tibia on the inner side (medial) of the leg and the fibula on the outer side (lateral) of the leg. The medial malleolus is the promine ...
has a triangular outline view showing a front concave surface.[
]
Phylogeny
''Morrosaurus'' was classified in the group Iguanodontia
Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', '' Dryosaurus'', '' Iguanodon'', '' Tenontosaurus'', and the hadrosaurids or "duck- ...
, as a basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
member of Euiguanodontia. This in turn formed a clade with other ornithopods of Patagonia
Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and ...
and Antarctica, particularly ''Trinisaura
''Trinisaura'' is an extinct genus of ornithopod dinosaur known from the lower levels of the Late Cretaceous Snow Hill Island Formation (early Maastrichtian stage) of James Ross Island, Antarctica. It contains a single species, ''Trinisau ...
'', '' Gasparinisaura'', '' Anabisetia'', '' Notohypsilophodon'', '' Talenkauen'' and '' Macrogryphosaurus'' in a group called Elasmaria
Elasmaria is a clade of ornithopods known from Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At ...
, whose members are distinguished by their adaptation to a running lifestyle which would be reflected by the narrow foot with a thin fourth metatarsal which indicates a high speed running; subsequently expanded chevrons, a feature that is associated with a greater surface area for attachment of the lateral muscles of the tail, which would give good control of the movements of this; and a curved humerus
The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
which demonstrates the absence of a deltopectoral ridge and therefore that the front leg was not used for walking. It cannot be determined, however, if ''Morrosaurus'' itself possessed these characteristics due to their limited remains. The existence of this clade may indicate that Patagonia, Antarctica and Australia shared the same type of fauna. The exact phylogenetic relationships within this clade could not be identified, except for ''Gasparinisaura'', which proved to be the most basal member of group.[
]Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
based in the phylogenetic analysis
In biology
Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that pro ...
of Rozadilla ''et al''., 2016:
See also
* South Polar region of the Cretaceous
The South Polar region of the Cretaceous comprised the continent of East Gondwana–modern day Australia and Antarctica–a product of the break-up of Gondwana in the Cretaceous Period. The southern region, during this time, was much warmer than i ...
* 2016 in paleontology
Flora Plants
Fungi
Cnidarians
Research
* '' Yunnanoascus haikouensis'', previously thought to be a member of Ctenophora, is reinterpreted as a crown-group medusozoan by Han ''et al.'' (2016).
* A study on the fossil corals from the Late T ...
References
{{Taxonbar, from=Q21122031
Ornithopods
Late Cretaceous dinosaurs
Maastrichtian life
Dinosaurs of Antarctica
Fossil taxa described in 2015
Taxa named by Fernando Novas
Ornithischian genera