''Morotopithecus'' is a genus of fossil ape discovered in

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

-age deposits of Moroto,

Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...

. The

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

status of ''Morotopithecus bishopi'' is debated to the extent that it challenges established views on the connection between Miocene primates and extant

hominids The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...

(i.e.

great apes The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...

Parsimonious In philosophy, Occam's razor (also spelled Ockham's razor or Ocham's razor; ) is the problem-solving principle that recommends searching for explanations constructed with the smallest possible set of elements. It is also known as the principle o ...

phylogenetic analyses indicate ''Morotopithecus'' is more derived than ''

Proconsul A proconsul was an official of ancient Rome who acted on behalf of a Roman consul, consul. A proconsul was typically a former consul. The term is also used in recent history for officials with delegated authority. In the Roman Republic, military ...

'', '' Afropithecus'', and '' Kenyapithecus'', but less derived than '' Oreopithecus'', '' Sivapithecus'', and '' Dryopithecus''. Under this arrangement, ''Morotopithecus'' would be a sister taxon to extant great apes while '' Hylobates'' (

gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical forests from eastern Bangladesh and Northeast Indi ...

s) seem to have branched off before this clade appeared. However, gibbons are believed to have branched off while ''Morotopithecus'' is dated to more than . In a comparison of teeth characteristics of ''Morotopithecus'' to '' Afropithecus'' the results showed little difference, plus evidence gathered from cranial comparisons also indicate that the two genera may be the same, a conclusion of limited confidence due to the lack of evidence to produce a complete anatomy for both (Patel, Grossman 2005). Meanwhile, Pickford (2002) referred the vertebrae to '' Ugandapithecus'', and considered ''Morotopithecus'' synonymous with ''Afropithecus''. It appears to have lived in wooded

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

, and had a diet dominated by leaves.

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Further reading