Moringaceae
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''Moringa'' is the sole
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
in the plant
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Moringaceae. It contains 13 species, which occur in
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
regions of Africa and Asia and that range in size from tiny herbs to massive trees. ''Moringa'' species grow quickly in many types of environments. The most widely cultivated species is '' Moringa oleifera'', native to the foothills of the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
in northwestern India, a
multipurpose tree Multipurpose trees or multifunctional trees are trees that are deliberately grown and managed for more than one output. They may supply food in the form of fruit, nuts, or leaves that can be used as a vegetable; while at the same time supplyi ...
cultivated throughout the tropics and marketed as a
dietary supplement A dietary supplement is a manufactured product intended to supplement a person's diet by taking a pill (pharmacy), pill, capsule (pharmacy), capsule, tablet (pharmacy), tablet, powder, or liquid. A supplement can provide nutrients eithe ...
, health food or source for herbalism practices. The fruit pods of ''Moringa oleifera'' ("drumsticks") are consumed as food in many parts of the world, particularly in
South Asia South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...
. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are commonly used to make tea. Oils are made from the seeds, while powders can be made from the leaves and
roots A root is the part of a plant, generally underground, that anchors the plant body, and absorbs and stores water and nutrients. Root or roots may also refer to: Art, entertainment, and media * ''The Root'' (magazine), an online magazine focusin ...
.


Description

''Moringa'' is considered one of the most widely diverse genera for its size ranging from small shrubs ('' M. pygmaea'') to large pachycaul trees ('' M. ovalifolia'').


Growth habit

''Moringa'' contains a wide range of growth
habits A habit (or wont, as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously. A 1903 paper in the ''American Journal of Psychology'' defined a "habit, from the standpoint of psychology, M. drouhardii, Moringa hildebrandtii">M. hildebrandtii,'' '' M. ovalifolia, M. stenopetala'' * Slender trees: '' M. concanensis, M. oleifera'', ''Moringa peregrina">M. peregrina'' * Moringa oleifera">M. oleifera'', ''Moringa peregrina">M. peregrina'' * M. arborea, M. ruspoliana'' * Moringa ruspoliana">M. ruspoliana'' * M. borziana, M. longituba, M. pygmaea,
M. rivae''


Leaves

Leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are typically pinnately compound with entire margins.


Flowers

Pinnation">pinnately compound with entire margins.


Flowers

bilaterally or Symmetry in biology">radially symmetric. Bottle trees typically produce small, radially symmetric flowers, while other members of the genus produce radially symmetric flowers. Most flowers range in color from white to cream to brown with the notable exception of ''M. longituba'' which produces bright red flowers.


Fruit

Fruits are typically elongated, slender, 3-valved "Pod (fruit), pods" resembling an Dehiscence (botany), indehiscent silique (in contrast with a true dehiscent silique). Fruits of ''M. oleifera'' ( Drumstick (fruit), drumstick), are a major agricultural product of
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, eaten as a
vegetable Vegetables are edible parts of plants that are consumed by humans or other animals as food. This original meaning is still commonly used, and is applied to plants collectively to refer to all edible plant matter, including edible flower, flo ...
and used for
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
.


Phytochemistry

''Moringa'' contain a number of sulfurous biochemical compounds called "mustard-oil glycosides" or glucosinolates commonly found in cruciferous vegetables of
Brassicaceae Brassicaceae () or (the older but equally valid) Cruciferae () is a medium-sized and economically important Family (biology), family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous pla ...
. Benzyl glucosinolate along with family-specific glucomoringin and glucosoonjnain have been detected from various ''Moringa'' species and are thought to be the cause of the bitter taste in some ''Moringa'' leaves.


Taxonomy


Higher-level classification

The
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
family, Moringaceae, containing genus ''Moringa'' has been placed in the order
Brassicales The Brassicales (or Cruciales) are an order (biology), order of flowering plants, belonging to the malvid group of eudicotyledons under the APG IV system. Well-known members of Brassicales include cabbage, cauliflower, Brussels sprout, broccoli, ...
according to most modern taxonomic systems, including the
APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publish ...
. Molecular data has suggested a close relationship between Moringaceae and Caricaceae with many identifying a "Caricaceae-Moringaceae" clade within Brassicales. Prior to the availability of molecular data, morphological classification of Moringaceae placed the family in either Brassicales or Sapindales due to the unusual morphological diversity of the family.


Classification within the genus

''Moringa'' contains three widely recognized clades—Donaldsonia, Moringa, and Dysmoringa. Donaldsonia, once thought to be a
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
of ''Moringa'', is a non-monophyletic clade identifiable by radially symmetric flowers and containing the bottle trees '' M. drouhardii, M. hildebrandtii,'' '' M. ovalifolia,'' and '' M. stenopetala.'' The Moringa clade contains all other members of genus ''Moringa'' (except ''M. longituba'') characterized by irregular floral symmetry, perigynous flowers, and short receptacles. The Dysmoringa clade contains the species ''M. longituba'' which diverges from common Moringa clade characteristics due to its long receptacle and red flowers. The exact phylogenetic relationship between members of ''Moringa'' continues to evolve with growing molecular data, though the Donaldsonia clade is consistently identified as the basalmost clade within the family.


List of species

* '' Moringa arborea'' Verdc. (indigenous to
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
) * '' Moringa borziana'' Mattei (indigenous to
Somalia Somalia, officially the Federal Republic of Somalia, is the easternmost country in continental Africa. The country is located in the Horn of Africa and is bordered by Ethiopia to the west, Djibouti to the northwest, Kenya to the southwest, th ...
) * ' Nimmo (indigenous to northern
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) * '' Moringa drouhardii'' Jum. – bottle tree (indigenous to southwestern
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
) * '' Moringa hildebrandtii''
Engl. Engl or Engl. may refer to: *England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more ...
– Hildebrandt's moringa (indigenous to southwestern Madagascar) * '' Moringa longituba'' Engl. (indigenous to
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
and Somalia) * '' Moringa oleifera'' Lam. (syn. ''M. pterygosperma'') – horseradish tree (indigenous to northwestern
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) * '' Moringa ovalifolia'' Dinter &
Berger Berger is a surname in both German language, German and French language, French, although there is no etymological connection between the names in the two languages. The French surname is an occupational name for a shepherd, from Old French ''bergi ...
(indigenous to
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
and
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
) * '' Moringa peregrina'' ( Forssk.) Fiori indigenous to Arabian Peninsula
Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004), ...
and in the Southern Sinai,
Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
* '' Moringa pygmaea'' Verdc. (indigenous to Somalia) * '' Moringa rivae'' Chiov. (indigenous to Kenya and Ethiopia) * '' Moringa ruspoliana'' Engl. (indigenous to Ethiopia) * '' Moringa stenopetala'' ( Baker f.) Cufod. (indigenous to Kenya and Ethiopia)


References


External links

* , Instituto de Biología, Universidad Nacional Autónoma de México
The International Moringa Germplasm Collection
{{Authority control Brassicales genera Edible plants