Montifringilla
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''Montifringilla'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds in the sparrow family
Passeridae Old World sparrows are a group of small passerine birds forming the family Passeridae. They are also known as true sparrows, a name also used for a particular genus of the family, '' Passer''. They are distinct from both the New World sparrows, ...
. It is one of three genera containing the snowfinches. As the English and scientific names suggest, these are high-altitude species, found in the mountain ranges of southern Eurasia, from the
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
east to the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
and western China.


Description

Snowfinches are stocky sparrows in length, with strong conical bills. They have pale brown upperparts, white underparts and extensive white panels in the wings, which transform them in flight. Adults may have black markings on the chin or around the eyes. Sexes are usually very similar, although the male white-winged snowfinch has a distinctive grey head. Young birds are a drabber version of the adult., , They have simple repetitive songs, given from a rock or during the elaborate circling display flight. The call is a simple ''chip'' or similar.


Ecology

Most snowfinches breed above altitudes of , but the white-winged snowfinch can occur from upwards. These hardy birds inhabit bare open mountain
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
. Snowfinches are not migratory but may move to lower altitudes or human habitation in winter, when these highly gregarious birds form large flocks. Snowfinches are primarily ground-feeding seed-eaters, though they also consume small
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, especially when breeding. They are typically fearless, and will forage around ski resorts, human habitation and rubbish tips. They nest in rock crevices, or more typically in holes left by
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s or (and even more often)
pika A pika ( , or ) is a small, mountain-dwelling mammal native to Asia and North America. With short limbs, a very round body, an even coat of fur, and no external tail, they resemble their close relative the rabbit, but with short, rounded ears. ...
s (Ochotonidae). The typical
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
is from three to six eggs.


Systematics and taxonomy

The genus ''Montifringilla'' was introduced by the German ornithologist
Christian Ludwig Brehm Christian Ludwig Brehm (24 January 1787 – 23 June 1864) was a German pastor and Ornithology, ornithologist. He was the father of the Zoology, zoologist Alfred Brehm. Life Brehm was born in Schönau (Odenwald), Schönau near Gotha on 24 Ja ...
in 1828. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was subsequently designated as the
white-winged snowfinch The white-winged snowfinch (''Montifringilla nivalis''), or snowfinch, is a small passerine bird. Despite its name, it is a sparrow rather than a true finch. Taxonomy In 1760 the French zoologist Mathurin Jacques Brisson included a description ...
. The name of the genus combines the Latin words ''mons'', ''montis'' "mountain" and ''fringilla'' "finch".


Species and systematics

The genus contains three
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
: Four species of snowfich are now separated in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Pyrgilauda ''Pyrgilauda'' is a genus of passerine birds in the sparrow family Passeridae. They are found in the Himalayas, Tibet and western China. The genus name was introduced by the French naturalist Charles Lucien Bonaparte in 1850. The type species wa ...
'', based on the genetics and the different vocalizations and
ecological Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...
preferences. Similarly, the white-rumped snowfinch is placed in a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus ''Onychostruthus''.


Taxonomy

The
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
s of ''Montifringilla'' have a convoluted history. ''Pyrgilauda'' is probably the most confused case among them: It was first established by
Charles Lucien Jules Laurent Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...
as a junior synonym of ''Pyrrhulauda'', and attributed to the brothers Edouard and
Jules Verreaux Jules Pierre Verreaux (24 August 1807 – 7 September 1873) was a French botanist and ornithologist and a professional collector of and trader in natural history specimens. He was the brother of Édouard Verreaux and nephew of Pierre Antoine Delal ...
. But ''Pyrrhulauda'' – established by Andrew Smith before Bonaparte – actually refers to certain sparrow-larks, today all placed in ''Eremopterix''. J. Verreaux indeed seems to have been the first to formally use ''Pyrgilauda'' for a particular snowfinch species, but that was only in 1871, when he described the small snowfinch. It was widely used for the southern snowfinch species in the late 19th and early 20th century, and continues to be used by some authors today. According to Article 11.6.1. and 50.7. in the third and fourth editions of the ''
International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted Convention (norm), convention in zoology that rules the formal scientific name, scientific naming of organisms treated as animals. It is also informally known as the I ...
'', the genus ''Pyrgilauda'' was thus validly established by Bonaparte as a possible snowfinch genus name – even though he did not intend to – because many authors after J. Verreaux used it for these birds, and not for the sparrow-larks. Bonaparte had correctly used ''Montifrigilla'' for the
white-winged snowfinch The white-winged snowfinch (''Montifringilla nivalis''), or snowfinch, is a small passerine bird. Despite its name, it is a sparrow rather than a true finch. Taxonomy In 1760 the French zoologist Mathurin Jacques Brisson included a description ...
– the only snowfinch known then – however, and thus ''Pyrgilauda'' had remained an unused "shell" of a
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
until someone placed a species in it. That was done by J. Verreaux, and thus many erroneously attributed authorship of the genus to him, although as per the ICZN ''Code'' Article 69.3. he technically just fixed its
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
as the small snowfinch. Another error occurred in 1982, when G. N. Kasin believed ''Pyrgilauda'' had been established twice, and thought the snowfinch genus name – which he believed to have been established by Verreaux in 1870 – was the
junior homonym In biology, a homonym is a name for a taxon that is identical in spelling to another such name, that belongs to a different taxon. The rule in the International Code of Zoological Nomenclature is that the first such name to be published is the s ...
of Bonaparte's sparrow-lark synonym. He set up ''Stepaniania'' to replace it. But with Bonaparte's name (as co-opted by Verreaux and later authors) being a valid junior subjective synonym and not preoccupied, ''Stepaniania'' itself becomes a junior objective synonym of ''Pyrgilauda''. The name ''Orospiza'' has a similar history to ''Pyrgilauda''. Bonaparte merely remarked it had been used by "some" (''aliquī'') unspecified authors, while many subsequent sources attributed it to
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...
. But that is a misunderstanding, and the only technically valid (though synonymous) name for the snowfinch genus proposed by Kaup was ''Chionospina''. Thus, in the case of ''Orospiza'' Bonaparte is again the valid author, though
Thomas Horsfield Thomas Horsfield (May 12, 1773 – July 24, 1859) was an American physician and natural history, naturalist who worked extensively in Indonesia, describing numerous species of plants and animals from the region. He was later a curator of the Eas ...
and
Frederic Moore Frederic Moore FZS (13 May 1830 – 10 May 1907) was a British entomologist and illustrator. He produced six volumes of ''Lepidoptera Indica'' and a catalogue of the birds in the collection of the East India Company. It has been said that Mo ...
were the first to set a type species. As ''M. nivalis'' was still the only known snowfinch in 1858 when they did this, ''Orospiza'' is a junior objective synonym of ''Montifringilla''. For somewhat different reasons, ''Orites'' is a junior objective synonym of ''Montifringilla''. First proposed for the
long-tailed tit The long-tailed tit (''Aegithalos caudatus''), also named long-tailed bushtit, is a common bird found throughout Europe and the Palearctic. The genus name ''Aegithalos'' was a term used by Aristotle for some European tits, including the long-tail ...
(''Aegithalos caudatus'') in Moehring's ''Geslachten der Vogelen'' in 1758, that widely ignored work was suppressed as a source of taxa by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
in 1987. Consequently, later attempts to re-establish that name as a junior synonym of '' Aegithalos'' are preempted by the erection of ''Orites'' for the white-winged snowfinch by
Eugen von Keyserling Eugen von Keyserling (22 March 1833 in Pockroy, Lithuania – 4 April 1889 in Dzierżoniów, Silesia Silesia (see names #Etymology, below) is a historical region of Central Europe that lies mostly within Poland, with small parts in the Czec ...
and Johann Heinrich Blasius in 1840. Meanwhile, Kaup's ''Chionospina'' was misspelled "Chionospiza" by
George Robert Gray George Robert Gray (8 July 1808 – 6 May 1872) was an English zoology, zoologist and author, and head of the Ornithology, ornithological section of the British Museum, now the Natural History Museum, London, Natural History Museum, London f ...
. As it, too, is a junior objective synonym of ''Montifringilla'',
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist, ornithologist and illustrator. It was he who first requested Leopold and Rudolf Blaschka, Leopold Blaschka to make a set of glass marine invertebrate m ...
's subsequent valid establishment of that name is still inconsequential – except for purely taxonomic purposes, as it makes neither Kaup nor Grey the valid author of ''Chionospiza'', but Reichenbach., , And finally, ''Onychostruthus'' had to be established because ''Onychospiza'',
Nikolai Przhevalsky Nikolay Mikhaylovich Przhevalsky (or Prjevalsky;; , . – ) was a Russian geographer and a renowned explorer of Central and East Asia. Although he never reached his ultimate goal, the city of Lhasa in Tibet, he still travelled through regio ...
's 1876 name for the proposed monotypic genus of the white-rumped snowfinch, was preoccupied. It had been given by Rey to the chestnut-eared bunting (today ''Emberiza fucata'') in 1872 already, as an unnecessary correction – and thus junior objective synonym – of Bonaparte's original ''Onychospina''.


Footnotes


References

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External links


Media related to Passeridae
on the Internet Bird Collection {{Taxonbar, from=Q922089   Taxa named by Christian Ludwig Brehm