''Mohanamico'' is an
extinct genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
New World monkey
New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboid ...
s from the
Middle Miocene (
Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the ...
in the
South American land mammal age
The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as age ...
s; 13.8 to 11.8 Ma). Its remains have been found at the
Konzentrat-Lagerstätte of
La Venta
La Venta is a pre-Columbian archaeological site of the Olmec civilization located in the present-day Mexican state of Tabasco. Some of the artifacts have been moved to the museum "Parque - Museo de La Venta", which is in nearby Villahermos ...
in the
Honda Group of
Colombia. The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''M. hershkovitzi''. Due to the relatively few material found of ''Mohanamico'', the placement of the genus is not certain and four possible families have been proposed by different authors,
Atelidae
The Atelidae are one of the five family (biology), families of New World monkeys now recognised. It was formerly included in the family Cebidae. Atelids are generally larger monkeys; the family includes the howler monkey, howler, spider monkey, ...
,
Callitrichidae
The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of th ...
,
Pitheciidae
The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region of B ...
or
Aotidae
Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South ...
.
Etymology
''Mohanimico hershkovitzi'' is named after the river god
Mohan of the
Magdalena River
The Magdalena River ( es, Río Magdalena, ; less commonly ) is the main river of Colombia, flowing northward about through the western half of the country. It takes its name from the biblical figure Mary Magdalene. It is navigable through much o ...
, in which valley the fossils were found and to honor
Philip Hershkovitz
Philip Hershkovitz (12 October 1909 – 15 February 1997) was an American mammalogist. Born in Pittsburgh, he attended the Universities of Pittsburgh and Michigan and lived in South America collecting mammals. In 1947, he was appointed a curator ...
for his contributions to the study of Colombian and other South American primates.
[Luchterhand et al., 1986, p.1757]
Description
Fossils of ''Mohanamico'' were discovered in the "Monkey Beds" of the
Honda Group, that has been dated to the
Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the ...
, about 12.5 Ma.
[Wheeler, 2010, p.133] ''Mohanamico'' was about the size of the living
squirrel monkey
Squirrel monkeys are New World monkeys of the genus ''Saimiri''. ''Saimiri'' is the only genus in the subfamily Saimirinae. The name of the genus is of Tupi origin (''sai-mirím'' or ''çai-mbirín'', with ''sai'' meaning 'monkey' and ''mirím'' ...
''
Saimiri sciureus
The Guianan squirrel monkey (''Saimiri sciureus'') is a species of squirrel monkey from Guiana, Venezuela and Brazil. ''S. sciureus'' formerly applied to Humboldt's squirrel monkey and Collins' squirrel monkey, but genetic research in 2009 an ...
''. Its
molars are low-crowned and the molar crests are not pronounced suggesting a frugivorous diet like ''
Aotus''. The lateral incisor is large and high-crowned, which foreshadows living
Pitheciidae
The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region of B ...
. The
canines
Canine may refer to:
Zoology and anatomy
* a dog-like Canid animal in the subfamily Caninae
** ''Canis'', a genus including dogs, wolves, coyotes, and jackals
** Dog, the domestic dog
* Canine tooth, in mammalian oral anatomy
People with the surn ...
and Pz were large and sharp like ''
Callimico
The Goeldi's marmoset or Goeldi's monkey (''Callimico goeldii'') is a small, South American New World monkey that lives in the upper Amazon basin region of Bolivia, Brazil, Colombia, and Peru. It is the only species classified in the genus ''Call ...
''. Analysis of the mandible and teeth suggest that ''Mohanamico'' is a primitive member of the Pitheciidae. Some similarities with ''Callimico'' and ''
Saguinus
The tamarins are squirrel-sized New World monkeys from the family Callitrichidae in the genus ''Saguinus''. They are the first offshoot in the Callitrichidae tree, and therefore are the sister group of a clade formed by the lion tamarins, Go ...
'' are also noted raising the possibility that pithecines and
callitrichids are monophyletic.
[Luchterhand et al., 1986, p.1753]
The estimated weight of ''Mohanamico'' was ,
[Silvestro, 2017, p.14] similar in size to ''
Aotus dindensis'' found in the same location.
[Gebo et al., 1990, p.738] The
Argentinian
Argentines (mistakenly translated Argentineans in the past; in Spanish ( masculine) or ( feminine)) are people identified with the country of Argentina. This connection may be residential, legal, historical or cultural. For most Argentines, ...
genus ''
Homunculus
A homunculus ( , , ; "little person") is a representation of a small human being, originally depicted as small statues made out of clay. Popularized in sixteenth-century alchemy and nineteenth-century fiction, it has historically referred to the ...
'' had smaller molars than ''Mohanamico''.
[Luchterhand et al., 1986, p.1756]
Some authors place ''Mohanamico'' in the
Callitrichidae
The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of th ...
,
[Tejedor, 2013, p.29] possibly related to the ''
Callimico
The Goeldi's marmoset or Goeldi's monkey (''Callimico goeldii'') is a small, South American New World monkey that lives in the upper Amazon basin region of Bolivia, Brazil, Colombia, and Peru. It is the only species classified in the genus ''Call ...
'' clade, based especially on its taller incisors and canines, large p2, and broader and longer trigonid in proportion to the talonid.
[Pérez et al., 2013, p.4] Other authors dispute this and consider the genus more likely a
Pitheciidae
The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region of B ...
,
[Wheeler, 2010, p.134] or possibly an
Aotidae
Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South ...
.
[Takai et al., 2001, p.303] It is also shown to be a stem
atelid.
[ If placed within the Pitheciidae, the primate would be at the base of the evolutionary radiation because of the large ]incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, w ...
s and the structure of its canines and premolar
The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth ...
s.[Defler, 2004, p.33]
Habitat
The Honda Group, and more precisely the "Monkey Beds", are the richest site for fossil primates in South America.[Rosenberger & Hartwig, 2001, p.3] It has been argued that the monkeys of the Honda Group were living in habitat that was in contact with the Amazon
Amazon most often refers to:
* Amazons, a tribe of female warriors in Greek mythology
* Amazon rainforest, a rainforest covering most of the Amazon basin
* Amazon River, in South America
* Amazon (company), an American multinational technolog ...
and Orinoco Basin
The Orinoco Basin is the part of South America drained by the Orinoco river and its tributaries. The Orinoco watershed covers an area of about 990000 km2, making it the third largest in South America, covering most of Venezuela and eastern p ...
s, and that La Venta itself was probably seasonally dry forest.[Lynch Alfaro et al., 2015, p.520] From the same level as where ''Mohanamico'' has been found, also fossils of ''Cebupithecia
''Cebupithecia'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of ...
'', ''Micodon
''Micodon'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of Col ...
'', '' Saimiri annectens'', ''Saimiri fieldsi
''Saimiri fieldsi'' is an extinct species of New World monkey in the genus '' Saimiri'' (squirrel monkeys) from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentra ...
'' and '' Stirtonia'' have been uncovered.[Setoguchi et al., 1986, p.762] From the same location, a fossil specimen of the fish tambaqui
The tambaqui (''Colossoma macropomum'') is a large species of freshwater fish in the family Serrasalmidae. It is native to tropical South America, but kept in aquaculture and introduced elsewhere. It is also known by the names black pacu, black ...
(''Colossoma macropomum'') was recovered.[
]
See also
* List of primates of Colombia
The primates of Colombia include 41 extant species in 13 genera and five families. Additionally, 12 fossil species in 10 genera and five families have been identified in Colombia, mainly at the La Venta Lagerstätte of the Honda Group, mostly from ...
* '' Canaanimico''
* ''Lagonimico
''Lagonimico'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of ...
''
* ''Patasola
The Patasola or "one leg" is one of many legends in South American folklore about female monsters from the jungle, appearing to male hunters or loggers in the middle of the wilderness when they think about women. The Patasola appears in the form ...
''
References
Bibliography
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Further reading
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{{Taxonbar, from=Q10331497
Prehistoric primate genera
Prehistoric monkeys
†Mohanamico
Miocene genus first appearances
Miocene extinctions
Miocene primates of South America
Laventan
Neogene Colombia
Fossils of Colombia
Honda Group, Colombia
Fossil taxa described in 1986