Mochlodon
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''Mochlodon'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of rhabdodontid
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
and
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
. It lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(85-80 Ma) and two
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are known: ''M. suessi'' and ''M. vorosi''.


Discovery and species

In 1859 coal mine administrator Pawlowitsch notified the
University of Vienna The University of Vienna (, ) is a public university, public research university in Vienna, Austria. Founded by Rudolf IV, Duke of Austria, Duke Rudolph IV in 1365, it is the oldest university in the German-speaking world and among the largest ...
that some fossils had been found in the ''Gute Hoffnung'' mine at Muthmannsdorf in
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
. A team headed by geologists
Eduard Suess Eduard Suess (; 20 August 1831 – 26 April 1914) was an Austrian geologist and an expert on the geography of the Alps. He is responsible for hypothesising two major former geographical features, the supercontinent Gondwana (proposed in 1861) and ...
and
Ferdinand Stoliczka Ferdinand Stoliczka (Czech language, Czech written Stolička, 7 June 1838 – 19 June 1874) was a Moravian palaeontologist who worked in India on paleontology, geology and various aspects of zoology, including ornithology, malacology, and herpe ...
subsequently uncovered numerous bones of several species, among them those of a euornithopod dinosaur. Stored at the university museum, the finds remained undescribed until they were studied by Emanuel Bunzel from 1870 onwards. Bunzel in 1871 named the euornithopod a new species of ''
Iguanodon ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species found worldwide have been classified in the genus ''Iguanodon'', dating from the Late Jurassic to Early Cretaceous, Taxonomy (bi ...
'': ''Iguanodon suessii''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours Suess and is today more often spelled ''suessi''. In 1881
Harry Govier Seeley Harry Govier Seeley (18 February 1839 – 8 January 1909) was a British paleontologist. Early life Seeley was born in London on 18 February 1839, the second son of Richard Hovill Seeley, a goldsmith, and his second wife Mary Govier. When his fa ...
named a separate genus: ''Mochlodon''. The generic name is derived from Greek ''mokhlos'', "bar", and ''odon'', "tooth", a reference to the bar-like median ridge on the teeth. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Mochlodon suessi''. ''Mochlodon'' and ''
Struthiosaurus ''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek ''sauros'' = lizard) is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.< ...
'', the latter found at the same site, are so far the only dinosaur genera named from Austrian finds. The
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
PIUW 2349 was found in the
Grünbach Formation The Grünbach Formation is an Austrian geological formation that dates to the early Campanian age of the Late Cretaceous. it forms part of the Gosau Group, and represents a marine regression event, representing a coastal/brackish environment, bei ...
of the Gosau Group dating from the Lower
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
, about 80 million years old. It consists of a
dentary In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone ...
, two
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e (presently lost), a parietal, a scapula, an ulna, a manual
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropod Sauropoda (), whose members are known as sauropods (; ...
, a femur and a tibia. Bunzel did not assign a
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
. In 2005 the dentary was chosen as the
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
. At the end of the nineteenth century Baron
Franz Nopcsa Franz may refer to: People * Franz (given name) * Franz (surname) Places * Franz (crater), a lunar crater * Franz, Ontario, a railway junction and unorganized town in Canada * Franz Lake, in the state of Washington, United States – see Fran ...
noted the similarity of fossils found in
Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
to both the French ''
Rhabdodon ''Rhabdodon'' (meaning "fluted tooth") is a genus of ornithopod dinosaur that lived in Europe approximately 70-66 million years ago in the Late Cretaceous. The genus contains a single species, ''R. priscus''. It is similar in build to a very rob ...
'' and the Austrian ''Mochlodon''. In 1899 he named some of these ''Mochlodon inkeyi'', the specific name honouring Béla Inkey, but the same year changed the name into ''Rhabdodon inkeyi''. In 1900 Nopcsa named some Romanian remains ''Mochlodon robustum'' (emended to ''M. robustus'' in 1990 by George Olshevsky). In 1915 however, he concluded that all this material could be referred to ''Rhabdodon'', the Austrian remains to ''Rhabdodon priscus''. In later years, ''Mochlodon'' was often considered a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''. In 2003, when ''M. robustus'' was renamed ''
Zalmoxes ''Zalmoxes'' is a genus of rhabdodontid ornithopod dinosaur from the Maastrichtian age of the Late Cretaceous in what is now Romania. The genus is known from specimens first named as the species '' Mochlodon robustum'' in 1899 by Franz Nopcsa ...
'', ''Mochlodon'' was tentatively reinstated as a separate genus for the species ''Mochlodon suessi''. In 2005 a study concluded that no unique derived features,
autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
, could be established for ''Mochlodon'' in relation to ''Zalmoxes'', assigning the Austrian remains provisionally to a ''Zalmoxes'' sp.; a definite identification would give ''Mochlodon'' nomenclatural priority. A second species, ''M. vorosi'', was described from the
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
aged
Csehbánya Formation The Csehbánya Formation is a Formation (geology), geological formation in the Transdanubian Mountains of Veszprém County, Hungary. The formation dates to the Late Santonian (around 85-84 million years ago) of the Late Cretaceous. It represents ...
of
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
in 2012.


Palaeoecology

Based on analysis of its tooth wear patterns, ''M. vorosi'' partitioned resources with the contemporary
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
'' Hungarosaurus'' by eating tougher vegetation than the latter, as its crowns wore down at more than twice the rate of the crowns of the
thyreophoran Thyreophora ("shield bearers", often known simply as "armored dinosaurs") is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous. Thyreophorans are characterized by the presence of bod ...
.


References

{{Taxonbar, from=Q18578509 Rhabdodontidae Dinosaur genera Santonian dinosaurs Campanian dinosaurs Fossil taxa described in 1881 Taxa named by Harry Seeley Dinosaurs of Europe