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The Miridae are a large and diverse insect family at one time known by the taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs".
Common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
s include plant bugs, leaf bugs, and grass bugs. It is the largest family of true bugs belonging to the suborder
Heteroptera The Heteroptera are a group of about 40,000 species of insects in the order Hemiptera. They are sometimes called "true bugs", though that name more commonly refers to the Hemiptera as a whole. "Typical bugs" might be used as a more unequivocal al ...
; it includes over 10,000 known species, and new ones are being described constantly. Most widely known mirids are species that are notorious agricultural pests that pierce plant tissues, feed on the sap, and sometimes transmit viral plant diseases. Some species however, are predatory.


Description

Miridae are small, terrestrial insects, usually oval-shaped or elongate and measuring less than in length. Many of them have a hunched look, because of the shape of the prothorax, which carries the head bent down. Some are brightly coloured and attractively patterned, others drab or dark, most being inconspicuous. Some genera are ant mimics at certain stages of life. The Miridae do not have any ocelli. Their rostrum has four segments. One useful feature in identifying members of the family is the presence of a cuneus; it is the triangular tip of the corium, the firm, horny part of the forewing, the hemelytron. The cuneus is visible in nearly all Miridae, and only in a few other
Hemiptera Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. ...
, notably the family Anthocoridae, which are not much like the Miridae in other ways. The tarsi almost always have three segments.


Some mirid species

* Lygus bugs (''Lygus'' spp.), including the tarnished and western tarnished plant bugs, are serious pests in the
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor p ...
,
strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely a ...
, and alfalfa industries. * ''
Stenotus binotatus ''Stenotus binotatus'' is a species of plant bug, originally from Europe, but now also established across North America and New Zealand. It is long, yellowish, with darker markings on the pronotum and forewings. It feeds on various grasses, an ...
'', a minor pest of cereal crops, especially
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeolog ...
* Apple dimpling bug (''
Campylomma ''Campylomma'' is a genus of bugs in the family Miridae and tribe Nasocorini.Henry TJ, Froeschner RC (1988). ''Catalog of the Heteroptera, True Bugs of Canada and the Continental United States''. Brill Academic Publishers. Species The following ...
liebknechti'') damages
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ances ...
blossoms and small growing fruits. * Mosquito bugs '' Helopeltis'' and ''
Afropeltis The genus ''Helopeltis'', also known as mosquito bugs, is a group of Heteropterans in the family Miridae (capsid bugs) and tribe Dicyphini. They include important pests of various crops, including cacao, cashew, cotton and tea. Now in a different ...
'' spp. – that infest various crops including
tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of '' Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of southwestern China and north ...
, cacao and
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor p ...
*
Honeylocust plant bug ''Blepharidopterus chlorionis'', known commonly as the honeylocust plant bug and under the synonym ''Diaphnocoris chlorionis'', is a hemipteran in the family Miridae. They are a major pest of the honey locust tree (''Gleditsia triacanthos''), a ...
(''Diaphnocoris chlorionis'') damages foliage on
honeylocust The honey locust (''Gleditsia triacanthos''), also known as the thorny locust or thorny honeylocust, is a deciduous tree in the family Fabaceae, native to central North America where it is mostly found in the moist soil of river valleys. Honey ...
trees. * Green mirid (''Creontiades dilutus'') damages many types of field crops. *
Potato capsid ''Closterotomus norvegicus'' (also known as the potato capsid) is a species of bugs belonging to the family Miridae, subfamily Mirinae. It can be found feeding on nettle, clover, and cannabis, as well as Compositae, potatoes, carrots and ...
(''Closterotomus norwegicus'') is a noted pest of
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Un ...
and
clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus h ...
plants in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
. * ''
Deraeocoris nebulosus ''Deraeocoris nebulosus'' is a species of plant bug in the family Miridae. It is found in North America. References Further reading * External links * Articles created by Qbugbot Insects described in 1872 Deraeocorini Hemipter ...
'' prefers other insects to plants in its diet, and has been used as a
biocontrol agent Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also inv ...
against
mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...
s and
scale insect Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than th ...
s. * '' Dicyphus hesperus'' sucks sap from various plants and preys on
whitefly Whiteflies are Hemipterans that typically feed on the undersides of plant leaves. They comprise the family Aleyrodidae, the only family in the superfamily Aleyrodoidea. More than 1550 species have been described. Description and taxonomy The ...
and red spider mites and can be used in biological pest control.


Systematics

This family includes a large number of
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, many of which are still unknown, distributed in more than 1300 genera. The taxonomic tree includes the following subfamilies and numerous tribes: * Bryocorinae **
Bryocorini ''Bryocorinae'' is a subfamily of bugs in the family Miridae. Tribes and genera ''BioLib''BioLib.cz
Bryocorinae Baerenspr ...
**
Dicyphini ''Dicyphini'' is a tribe of bugs in the family Miridae. Subtribes and Genera The ''On-line Systematic Catalogue of Plant Bugs'' includes: Subtribe Dicyphina * '' Campyloneura'' Fieber, 1861 * '' Campyloneuropsis'' Poppius, 1914 * '' Chius'' * ...
**
Eccritotarsini Eccritotarsini is a tribe of plant bugs in the family Miridae. There are about 14 genera and at least 40 described species in Eccritotarsini. Genera These 14 genera belong to the tribe Eccritotarsini: * '' Aguayomiris'' Maldonado Caprilles, 1 ...
* Cylapinae ** Cylapini ** Fulviini * Deraeocorinae ** Clivinematini **
Deraeocorini Deraeocorini is a tribe of plant bugs in the family Miridae. There are at least 19 genera and more than 340 described species in Deraeocorini. Genera These 19 genera belong to the tribe Deraeocorini: * '' Acutifromiris'' Hernandez & Stonedahl ...
** Hyaliodini ** Saturniomirini ** Surinamellini ** Termatophylini * Isometopinae ** Diphlebini ** Isometopini *
Mirinae Mirinae is a subfamily of plant bugs, insects in the family Miridae. Tribes * Herdoniini * Hyalopeplini * Mecistoscelini * Mirini * Restheniini * Scutelliferini * Stenodemini Stenodemini is a tribe of plant bugs in the family Miridae. ...
**
Herdoniini Herdoniini is a tribe of plant bugs in the family Miridae The Miridae are a large and diverse insect family at one time known by the taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs". Common ...
** Hyalopeplini **
Mirini Mirini is a tribe of plant bugs belonging to the subfamily Mirinae. Genera '' Acanthocranella'' - '' Acanthopeplus'' - '' Actinonotus'' - '' Adelphocoridea'' - ''Adelphocoris'' - '' Adelphocorisella'' - '' Adnotholopus'' - '' Adphytocoris'' - ...
** Pithanini **
Restheniini Restheniini is a tribe of plant bugs in the family Miridae. There are at least 4 genera and 30 described species in Restheniini. Genera These four genera belong to the tribe Restheniini: * ''Oncerometopus'' Reuter, 1876 * '' Opistheurista'' Car ...
**
Stenodemini Stenodemini is a tribe of plant bugs in the family Miridae. There are more than 60 described species in Stenodemini. Genera ''BioLib'' includes: # '' Acetropis'' Fieber, 1858 # '' Acomocera'' Eyles, 1975 # ''Actinocoris'' Reuter, 1878 # '' Ast ...
* Orthotylinae ** Ceratocapsini ** Halticini ** Orthotylini *
Phylinae Phylinae is a subfamily of the plant bug family Miridae. Species of this family are found worldwide. In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, an ...
**
Hallodapini Hallodapini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Hallodapini. Most species of this tribe are found in the old world, including Australia. Only one genus, Cyrtopeltocoris, is found in North America. Ge ...
**
Leucophoropterini Leucophoropterini is a tribe of plant bugs in the family Miridae. There are more than 20 genera and over 100 species in Leucophoropterini. Most species of this tribe are found in eastern Asia and Australia, with a single genus, ''Tuxedo'', found ...
**
Phylini Phylini is a tribe of plant bugs in the family Miridae, based on the type genus ''Phylus''. There are at least 440 described species in Phylini. Subtribes and selected Genera * Full list of Phylini genera here Keltoniina Auth. Schuh & Menard, ...
**
Pilophorini Pilophorini is a tribe of plant bug. The type genus is ''Pilophorus'' (Hahn, 1826). Schuh's analysis indicates that the Pilophorini originated in tropical Gondwanaland and subsequently spread into the temperate Northern Hemisphere, where they ...


Psallopinae

Auth.: Schuh, 1976 # '' Isometopsallops'' Herczek & Popov, 1992 # '' Psallops'' Usinger, 1946 # †'' Cylapopsallops'' Popov & Herczek, 2006 # †'' Epigonopsallops'' Herczek & Popov, 2009


Genera ''

Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''

''BioLib'' includes: # '' Amulacoris'' Carvalho & China, 1959 # '' Anniessa'' Kirkaldy, 1903 # '' Auchus'' Distant, 1893 # '' Bahiarmiris'' Carvalho, 1977 # '' Brasiliocarnus'' Kerzhner & Schuh, 1995 # '' Carmelinus'' Carvalho & Gomes, 1972 # '' Carmelus'' Drake & Harris, 1932 # '' Chaetophylidea'' Knight, 1968 # '' Charitides'' Kerzhner, 1962 # '' Colimacoris'' Schaffner & Carvalho, 1985 # '' Cylapocerus'' Carvalho & Fontes, 1968 # '' Duckecylapus'' Carvalho, 1982 # '' Englemania'' Carvalho, 1985 # '' Eurycipitia'' Reuter, 1905 # '' Faliscomiris'' Kerzhner & Schuh, 1998 # '' Fuscus'' Distant, 1884 # '' Guerrerocoris'' Carvalho & China, 1959 # ''Gunhadya'' - monotypic '' Gunhadya rubrofasciata'' Distant, 1920 # '' Heterocoris'' Guérin-Ménéville in Sagra, 1857 # '' Knightocoris'' Carvalho & China, 1951 # '' Leonomiris'' Kerzhner & Schuh, 1998 # '' Macrotyloides'' Van Duzee, 1916 # '' Merinocapsus'' Knight, 1968 # '' Mircarvalhoia'' Kerzhner & Schuh, 1998 # '' Montagneria'' Akingbohungbe, 1978 # '' Muirmiris'' Carvalho, 1983 # '' Myochroocoris'' Reuter, 1909 # '' Nesosylphas'' Kirkaldy, 1908 # '' Notolobus'' Reuter, 1908 # '' Nymannus'' Distant, 1904 # '' Paracoriscus'' Kerzhner & Schuh, 1998 # '' Paraguayna'' Carvalho, 1986 # '' Prodomopsis'' TBD # '' Prodomus'' TBD # '' Pseudobryocoris'' Distant, 1884 # '' Pygophorisca'' Carvalho & Wallerstein, 1978 # '' Rayeria'' TBD # '' Rewafulvia'' Carvalho, 1972 # '' Rhynacloa'' Reuter # '' Rondonisca'' Carvalho & Costa, 1994 # '' Rondonoides'' Carvalho & Costa, 1994 # '' Rondonotylus'' Carvalho & Costa, 1994 # '' Spanogonicus'' Berg # '' Sthenaridia'' TBD # ''
Zoilus Zoilus ( el, Ζωΐλος ''Zoilos''; c. 400320 BC) was a Greek grammarian, Cynic philosopher, and literary critic from Amphipolis in Eastern Macedonia, then known as Thrace. He took the name Homeromastix (Ὁμηρομάστιξ "Homer whippe ...
'' Distant, 1884


References


Further reading

* *
Google books preview


External links


PBI Plant Bug Index

Link to Al Wheeler's book on the biology of mirids

Research on mirids of Southern California at UC Riverside


on the
University of Florida The University of Florida (Florida or UF) is a public land-grant research university in Gainesville, Florida. It is a senior member of the State University System of Florida, traces its origins to 1853, and has operated continuously on its ...
/
Institute of Food and Agricultural Sciences The University of Florida Institute of Food and Agricultural Sciences (UF/IFAS) is a teaching, research and Extension scientific organization focused on agriculture and natural resources. It is a partnership of federal, state, and county governm ...
''Featured Creatures'' website {{Taxonbar, from=Q864705 Heteroptera families Articles containing video clips Taxa named by Carl Wilhelm Hahn